• Agius C. 1981. Preliminary studies on the ontogeny of the melanomacrophages of teleost hematopoetic tissues and age-related changes. Dev Comp Immunol 5: 597606.
  • Agius C, Agbede SA. 1984. An electron microscopical study on the genesis of lipofuscin, melanin and haemosiderin in the haemopoietic tissues of fish. J Fish Biol 24: 471488.
  • Agius C, Roberts RJ. 2003. Review: melano-macrophage centres and their role in fish patology. J Fish Biol 26: 499509.
  • Bagnara J, Matsumoto J. 2006. Comparative anatomy and physiology of pigment cells in nonmammalian tissues. In: Nordlund JJ, Boissy RE, Hearing VJ, King RA, Ortonne JP, editors. The pigmentary system: physiology and pathophysiology. New York: Oxford University Press. p 940.
  • Barni S, Bertone V, Croce AC, Bottiroli G, Bernini F, Gerzeli G. 1999. Increase in liver pigmentation during natural hibernation in some amphibians. J Anat 195: 1925.
  • Fenoglio C, Boncompagni E, Fasola M, Gandini C, Comizzoli S, Milanesi G, Barni S. 2005. Effects of environmental pollution on the liver parenchymal cells and Kupffer-melanomacrophagic cells of the frog Rana esculenta. Ecotoxicol Environ Saf 60: 259268.
  • Franco-Belussi L, Oliviera C. Lippopolysacharides induces changes in the visceral pigmentation of Eupemphix nattereri (Anura: Leiuperidae). Zoology 114:298305.
  • Franco-Belussi L, Santos LRS, Zieri R, Oliveira C. 2011. Visceral pigmentation in Dendropsophus (Anura: Hylidae): occurrence and comparison. Zool Anz 250: 102110.
  • Franco-Belussi L, Zieri R, Santos LRS, Moresco RM, Oliveira C. 2009. Pigmentation in anuran testes: anatomical pattern and variation. Anat Rec 292: 178182.
  • Gallone A, Guida G, Maida I, Cicero R, 2002. Spleen and liver pigmented macrophages of Rana esculenta L. A new melanogenic system? Pigment Cell Res 15: 3240.
  • McGraw KJ, 2005. The antioxidant function of many animal pigments: are there consistent health benefits of sexually selected colourants? Rev Anim Behav 69: 757764.
  • Moresco RM, Oliveira C. 2009. A comparative study of the extracutaneous pigmentary system in three anuran amphibian species evaluated during the breeding season. S Am J Herpetol 4: 18.
  • Oliveira C, Zieri R. 2005. Pigmentação testicular em Physalaemus nattereri (Steindachner) (Amphibia, Anura) com observações anatômicas sobre o sistema pigmentar extracutâneo. Rev Bras Zool 22: 454460.
  • R Development Core Team. 2010. R: a language and environment for statistical computing. R Foundation for Statistical Computing, Vienna, Austria. Available at: http://www.R– Accessed on 20 June 2010.
  • Sichel G, Scalia M, Mondio F, Corsaro C, 1997. The amphibian Kupffer cells build and demolish melanosomes: an ultrastructural point of view. Pigment Cell Res 10: 271287.
  • Sokal RR, Rohlf FJ. 1995. Biometry: the principles and practice of statistics in biological research. 3rd ed. New York: W.H. Freeman and Company.
  • Zieri R, Taboga SR, Oliveira C. 2007. Melanocytes in the testes of Eupemphix nattereri (Anura, Leiuperidae): histological, estereological and ultrastructural aspects. Anat Rec 290: 795800.
  • Zuasti A, Jiménez-Cervantes C, Gargia-Borrón JC, Ferrer C. 1998. The melanogenic system of Xenopus laevis. Arch Histol Cytol 61: 305316.