Previous work on stream channels in upland areas of Britain has demonstrated a close control over channel morphology and stability by the rate of coarse sediment supply from the hillslopes of the catchment. Streams fed by large amounts of coarse sediment develop unstable, wide, often braided channels, whereas those with limited coarse sediment supply develop stable, much narrower, often meandering channels. The sediment supply from hillslopes is controlled by thresholds of hillslope stability, storm event frequency, and the coupling between the hillslopes and the channel. Climatically-induced changes in any of these three factors may have implications for channel morphology and stability. This paper examines these implications in British upland fluvial systems, with particular reference to the Howgill Fells, Cumbria, in the contexts of the adjustment of stream channels to sediment supply from erosional gully systems, and their response to and recovery from major flood events.