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 The quantification of inter-ocean leakage from the South Indian to the South Atlantic Ocean is an important measure for the role of the Agulhas system in the global thermohaline circulation. To explore the specific role of mesoscale variability (such as Agulhas rings and Mozambique eddies) in this process a high-resolution model (based on NEMO-ORCA) for the Agulhas region has been set up. It is nested into a global coarse-resolution model. The high-resolution nest captures all salient features of the greater Agulhas region, including the upstream perturbations of the Agulhas Current and Natal Pulses along the African coast. A comparison of the inter-ocean exchange in the high-resolution nest with its coarse resolution counterpart reveals that the latter significantly over-estimates the amount of water flowing into the Atlantic Ocean, demonstrating the need to explicitly simulate the mesoscale features. A sensitivity experiment that excludes the upstream perturbations revealed no difference in the amount of inter-ocean exchange. However, the realistic representation of Agulhas rings and their drift path into the South Atlantic depends on the simulation of those upstream perturbations.
 The global thermo-haline circulation of the ocean has to pass through a few chokepoints [Gordon, 1986] of which the region directly south of Africa is one. Here warm and salty subtropical water from the Indian Ocean flows into the South Atlantic, providing about two-thirds of the upper limb of the meridional overturning circulation in the Atlantic [Weijer et al., 1999]. The mechanisms for this leakage consist largely of the shedding of Agulhas rings at the Agulhas Current's retroflection [Lutjeharms, 2006], an intermittent process apparently affected by perturbations originating in the upstream portions of the Agulhas Current [van Leeuwen et al., 2000]. The generation and role of these mesoscale processes in the production of Agulhas rings, and their effect on the net transport of water from the Indian to the Atlantic Ocean, is studied by utilizing a novel, high-resolution “two-way nesting” model of the Agulhas regime. It provides a realistic simulation of the mesoscale processes triggering the shedding and propagation of rings, including the emergence of eddies in the Mozambique Channel that are found critical for the generation of Natal Pulses [Schouten et al., 2002], and thus, upstream retroflection events of the Current. The sequence of model experiments demonstrates the effective control of the net inter-ocean exchange south of Africa by mesoscale dynamics.
 The Agulhas Current is the major western boundary current of the southern hemisphere with an average volume flux [Beal and Bryden, 1999] of about 70 Sv (1 Sv = 1 × 106 m3s−1). Shortly after passing the southern tip of the African continent, the bulk of the current turns east in the Agulhas Retroflection, with only about 20% of the volume flux leaking into the South Atlantic [de Ruijter et al., 1999a]. Much of the leakage is associated with the shedding of Agulhas rings at the retroflection that are the biggest anti-cyclonic vortices observed in the world ocean [Olson and Evans, 1986]. The importance of this exchange is emphasized by paleo-oceanographic studies [Peeters et al., 2004] which suggest that the Indian to Atlantic Ocean connection has undergone large oscillations during glacial cycles, apparently linked to variations in upper-layer temperatures of the South Atlantic, and the deep water formation in the sub-polar North Atlantic. Accordingly, lack of a realistic Agulhas retroflection has been identified as a critical deficiency of global climate models, with salinity anomalies due to false inflows of Indian Ocean water being advected throughout the Atlantic and impacting its meridional overturning circulation [Banks et al., 2007, Weijer et al., 1999]. A realistic simulation of the interocean exchange is a formidable challenge due to the highly nonlinear dynamics of the retroflection regime, including its little known dependence on mesoscale flow perturbations further upstream. Present understanding of these perturbations and their effect on the retroflection can be summarized as follows.
 The trajectory of the Agulhas Current's northern part is very stable [Gründlingh, 1983]. This path invariance is interrupted by the intermittent passage of a singular meander, the Natal Pulse [Lutjeharms and Roberts, 1988], proceeding from a region of reduced gradient in the shelf slope at the Natal Bight (Figure 1). It has been shown [de Ruijter et al., 1999b] on theoretical grounds that this is the only location along the path of the northern Agulhas Current where the bathymetry will allow instabilities in the trajectory to develop. This can only come about if the current intensity exceeds a certain threshold and it has been suggested [de Ruijter et al., 1999b] that this could be achieved by the adsorption of an anti-cyclonic eddy onto the current. To date only one case study [Schouten et al., 2002] has actually shown this to occur. There is evidence [van Leeuwen et al., 2000] that the downstream movement of Natal Pulses may cause the shedding of Agulhas rings at the retroflection. Natal Pulses have also been implicated in the upstream retroflection of the Agulhas Current [Lutjeharms and van Ballegooyen, 1988] through which all the water in the Agulhas Current is siphoned off to the Agulhas Return Current at an early stage with no normal inter-ocean exchange being feasible for a while.
 Previous modelling studies have begun to capture these intricacies of the Agulhas system, however, at the expense of the modelled frequency and dimensions of rings being shed in the retroflection region due to compromises between resolution [Biastoch and Krauss, 1999] and domain sizes [Speich et al., 2006]. In this study, the question how important these mesoscale processes are for the net interocean exchange south of Africa is addressed by a sequence of experiments with a global ocean – sea-ice model at 1/2° resolution (ORCA05) based on the NEMO code [Madec, 2006]. It is forced by bulk formulae using daily, inter-annually varying wind and thermohaline surface forcing fields over the period 1958–2004 [Large and Yeager, 2004]. Its main, novel element is the “two-way nesting” [Debreu et al., 2008] of a high-resolution, 1/10° model (AG01-R) of the Agulhas region (20°W–70°E, 47°S–7°S, red box in Figure 1) into this coarser, well established global model [Biastoch et al., 2008]. The nesting approach not only allows one to study the effect of outside perturbations on the Agulhas regime, but also the feedback of mesoscale Agulhas dynamics on the large-scale circulation, a characteristic that is not feasible with global high-resolution models [Maltrud and McClean, 2005]. The model features partially filled bottom cells (46 levels) and advanced advection schemes which have been shown to be crucial elements of a reasonable simulation of the Agulhas regime [Barnier et al., 2006]. In a second set-up of the nested model (AG01-S), the specific role of Mozambique eddies is examined by placing the northern boundary of the nest at 27°S, leaving the region around Madagascar at coarse resolution.
3. Representation of the Inter-ocean Exchange
 The main experiment (AG01-R) succeeds in representing the salient features of the Agulhas regime in a highly realistic way (Figure 1; see also auxiliary material). These include the Agulhas Current with a realistic transport of 71 ± 16.4 Sv at 32°S (concurring with estimates based on hydrographic sections [Beal and Bryden, 1999]) characteristically hugging the shelf edge and extending 200 km offshore; the highly variable Agulhas Undercurrent of 4.8 ± 3.8 Sv [Bryden et al., 2005]; Agulhas rings with a diameter of 360 ± 40 km moving off into the South Atlantic with a translation speed of about 24 km/week and an Agulhas Return Current [de Ruijter et al., 1999a] with a realistic representation of its average location, variability and shedding of eddies to either side. About 5 ± 1 anti-cyclonic Mozambique eddies, matching observations in size and frequency [Schouten et al., 2002], are formed per year with average diameters of 270 ± 45 km and extending to the sea floor. Natal Pulses are evident on the onshore side of the Agulhas Current at least 2 ± 1 times per year, moving downstream with an average rate of about 22 km/day. Upstream retroflections [Lutjeharms and van Ballegooyen, 1988] are seen in the expected location south of Port Elizabeth at 26 ± 1°E. What is the role of the host of mesoscale processes interacting with the Agulhas Current in the net volume transfer between the Indian and Atlantic Ocean? This question is addressed by comparing the reference simulation (AG01-R) with an experiment in which the same global model was integrated without the high-resolution nest in the Agulhas regime. The solution of this non-eddying model (ORCA05) portrays the inter-oceanic exchange as a continuation of parts of the Agulhas Current as a smooth current, reminiscent of coarse resolution models typically used for climate studies. For assessing the portion of the Agulhas water flowing into the South Atlantic the Eulerian model field does not provide an optimal measure since mesoscale eddies lead to enormous fluctuations in the east- and westward velocities involving a complex suite of time-dependent recirculation features. We therefore adopted a Lagrangian diagnostic [Blanke et al., 1999] based on tracking artificial “fluid” particles, where a large number of particles were seeded continuously over a time span of 4 years into the modelled Agulhas Current at 32°S. Each particle represents a certain amount (max. 0.01 Sv) of the total transport and is advected by the time-dependent flow field. At specified control sections in the Atlantic and Indian Ocean (Figure 2) particles are counted to give an average transport, similar to the analysis in the regional model by Speich et al. . In the reference simulation (AG01-R), with the eddy-resolving Agulhas nest, about 12 Sv (about one fifth of the Agulhas transport of 63 Sv at 32°S) reaches the northern/western sections in the Atlantic, representing the portion of upper water masses of the inter-oceanic exchange south of Africa, consistent with the range of observational estimations [de Ruijter et al., 1999a]. This is in marked contrast to the non-eddying case (ORCA05), where a much higher fraction (one third) of the Agulhas water enters the Atlantic. (According to the theoretical study of Dijkstra and de Ruijter , the leakage in this non-eddying case may be dependent by model parameters controlling the inertial-viscous character of the boundary layer regime.) The pronounced contrast between the two simulations with identical global conditions demonstrates that the supply of Indian Ocean water to the South Atlantic, and thus, the warm water path of the global thermohaline circulation, is not determined by large-scale dynamics alone, but significantly influenced by the regional, mesoscale dynamics of the Agulhas regime.
4. Role of Upstream Peturbations
 Upstream retroflections (examples for regular and upstream retroflection states are given in Figure 3) of the Agulhas Current are a prominent feature of the Agulhas dynamics, effectively causing one of the largest western boundary current in the world ocean to short-cut its south-western path for about 2–3 months. It has been assumed that these upstream retroflections are forced by passing Natal Pulses [Lutjeharms and van Ballegooyen, 1988] and that they may lead to substantial reductions in the inter-oceanic exchange [Lutjeharms and de Ruijter, 1996]. The time evolution of a coastal section along the African continent (Figure 3) indicates that the 5–6 Mozambique eddies generated per year are a necessary condition for the generation of Natal Pulses; however, this is not a sufficient condition since only 2–5 Natal Pulses per year appear in the same period. Furthermore, only large Natal Pulses, causing a displacement of the core of the Agulhas Current by more than 400 km offshore, in combination with a northward extension of the meander of the Agulhas Return Current are able to cause upstream retroflections. Such instances are rare (about 1–2 per year in our example). Similar numbers can be gathered from observations (here not shown), although a one-by-one comparison is difficult to perform due to limitations in the observational data base and would require a dedicated along-track study [Schouten et al., 2002]. Figure 3b shows the almost complete absence of Natal Pulses in a sensitivity experiment without Mozambique eddies (AG01-S). Some single Pulses (e.g., in 1971) were generated by eddies from further offshore, but these are too weak and therefore do not cause any upstream retroflections. However, the modelled upstream retroflections are found to have a negligible effect on net inter-ocean exchange (Figure 2), thus not supporting previous hypotheses arguing for a large-scale relevance of these events [Lutjeharms and de Ruijter, 1996].
 The independency of the magnitude of inter-oceanic volume flux on upstream retroflections does not imply that mesoscale upstream control mechanisms are unimportant for a proper description of the Agulhas leakage. Although simulating a similar number of Agulhas rings, those in the sensitivity experiment without Mozambique eddies show uncharacteristic regularity in time and space. As a consequence the ring paths are much more constrained in the South Atlantic, like “pearls on a string” (Figure 4); a behaviour that is typical for coarser models [Barnier et al., 2006] and also found in limited-area simulations [Speich et al., 2006]. It is unclear whether a false regularity in the ring spreading may influence the temporal variability of Agulhas leakage or might bias water masses in the South Atlantic where Agulhas rings finally end up and release their anomalous loads of heat and salt. In a similar modelling study Penven et al.  also found the interoceanic transport to be insensitive to upstream perturbations (although they did not report differences in ring paths, probably because of the coarser resolution in the Agulhas retroflection area or the weak but still existent upstream perturbations). However, the uniform eddy-permitting resolution of their model did not allow isolating the mesoscale in the Agulhas retroflection area itself.
 Based on a realistic portrayal of the salient features of the Agulhas system, the model configuration advances our process understanding of the Agulhas system and demonstrates the significance of the local, mesoscale flow dynamics for the inter-ocean exchange south of Africa. Critical elements include the triggering of Natal Pulses by offshore eddies, the spawning of Agulhas rings with or without the advent of Natal Pulses, and the significant dependence of upstream retroflection on Natal Pulses. A reliable representation of these processes appears of paramount importance for assessing the role of this regime in model simulations of past and future climate changes [Peeters et al., 2004].
 The integration of the experiments has been performed at the Höchstleistungsrechenzentrum Stuttgart (HLRS) and the Computing Centre at Kiel University. We gratefully thank the NEMO, AGRIF and ARIANE System Teams as well as Jean-Marc Molines for their technical support. The analysis was performed under the DFG project BO 907/2-2. J.L. received support from the Alexander von Humboldt-Stiftung.