We explore the extent to which it is possible to convert erosion rate data into uplift rate or erosion laws, using a landscape evolution model. Transient stages of topography and erosion rates of a block uplifting at a constant rate are investigated at different spatial scales, for a constant climate, and for various erosion laws and initial topographies. We identify three main model types for the evolution of the mountain-scale mean erosion rate: “linear”-type, “sigmoid”-type and “exponential”-type. Linear-type models are obtained for topographies without drainage system reorganization, in which river incision rates never exceed the uplift rate and stepped river terraces converge upstream. In sigmoid-type and exponential-type models (typically detachment-limited or transport-limited models with a significant transport threshold), drainage growth lasts a long time, and correspond to more than linear transport laws in water discharge and slope. In exponential-type models, the mean erosion rate passes through a maximum that is higher than the rock uplift rate. This happens when the time taken to connect the drainage network exceeds half the total response time to reach dynamic equilibrium. River incision rates can be much greater than the uplift rate in both cases. In the exponential-type model, river terraces converge downstream. Observations of a mountain in the Gobi-Altay range in Mongolia support the exponential-type model. This suggests that the erosion of this mountain is either detachment-limited or transport-limited with a significant transport threshold. This study shows that drainage growth could explain differences in erosion rate measurements on different spatial scales in a catchment.
 In this paper, we address the following question: what controls the shape of an ε versus time curve? We explore the hypothesis that (1) erosion rates on a steadily rising block may following one of a small number of characteristic patterns, and (2) these patterns involve systematic changes over time in the relationship between erosion rate and rock uplift rate. Analyzing these changes, we evaluate if it is possible to extract information about uplift rate and erosion laws from topographic and erosion rate data on different spatial scales from a catchment in a transient state. We investigate the relationships between the mountain-scale erosion rate, local-scale river incision rate, and river transient profiles in the case of an uplifting surface with a constant area and subject to constant and homogeneous uplift and climate. Restricting the study to a constant climate is for the sake of simplicity. Thus, erosion rates are calculated over timescales that express average global climate variations.
 We use a landscape evolution model (CIDRE), which allows diffusive and advection transport to be modeled over geological time spans. We analyze the effect of erosion laws and the initial topography on the evolution of topography and erosion rates.
diffusion sediment capacity per unit width toward cell j
transport capacity per unit width toward cell j
density of water
critical slope for material collapse (0.6 for sediment and 1.7 for bedrock)
critical shear stress
uplift rate (1 mm a−1)
flow width in the direction toward cell j
water fall velocity of clast in still water
 Hydrology is modeled by propagating a specified volume of water per unit of time downstream from the highest to the lowest cell, in order to ensure water conservation in the drainage network. For each cell, the water flux Q [L3T−1] is the sum of all incoming water fluxes from upstream draining cells, as well as the local precipitation:
where Δx [L] is the cell width, Pi and P [LT−1] are the rates of precipitation over the upstream cells i, and the local precipitation rate, respectively. In holes, water disappears entirely for the purpose of the present study. Neglecting water accumulation in lakes can have potential impact on our results, which we discuss in section 5.2.
in which Sj is the slope toward a cell j. The number of cells j depends on the topography. It is usually 3 or 4 on hillslopes and is generally reduced to one in a river network, as with the “steepest-descent” algorithm.
 Processes involving small transport distances, such as soil creep and small landslides, are modeled using a transport rate per unit width qdj [L2T−1] toward each lower neighboring cell j, which is assumed to depend nonlinearly on the local elevation gradient [Roering et al., 1999]:
where κ [L2T−1] is a diffusion coefficient, Sc is the critical slope corresponding to the material gradient of repose. In the case of bedrock, the use of this law assumes that the weathering rate is sufficient to erode the bedrock at the rate determined by κ, and that bedrock collapses for slopes approaching Sc. A linear approximation of this law is used for slopes greater than 0.95Sc to prevent infinite flux.
 Sediment and bedrock layers have distinct lithological characteristics. The transport rate of sediments carried by water is determined by the specific transport capacity expressed as a sediment flux per unit width qtj [L2T−1]
where Wj is the flow width [L] in each lower direction j, and τc is a transport threshold, while other parameters are positive constants. The flow width Wj [L] in each lower direction j is assumed to scale with the corresponding water flux volume:
where kw is a constant.
Equation (4) can be viewed as a development of the classic form of the excess shear stress formula (qtj = Kall(τ − τc)p), assuming uniform flow conditions, Manning or Chezy friction laws and a small depth to width hydraulic geometry [Tucker, 2004]. In this case, τc [ML−1T−2] is a critical shear stress for clast entrainment, kt, m and n are linked to the Manning or Chezy friction parameters, and Kall is an alluvial transport coefficient (see, e.g., Tucker  for details).
with α, β and a as positive constants and Kbr representing a bedrock detachment coefficient. The detachment capacity can be viewed as deriving from the excess shear stress formula (Ij = Kbr (τ − τc)a), in which case α = m and β = n. The same critical shear stress τc is used for both bedrock detachment and sediment entrainment, although they can be set differently in the model.
 It is important to note that the dimension of Kall and Kbr depends on p and a, which changes the meaning of these constants from one experiment to another and thus makes comparison difficult. It is easier to compare if the transport capacity and bedrock incision rates are nondimensionalized, as in Appendix A.
 In order to represent the fact that carried sediment can enhance (“tool effect”) or inhibit (“cover effect”) bedrock erosion, the following incision law can be used:
This is a development of the Sklar and Dietrich's  saltation-abrasion model, using the same assumptions as in equation (4). Qs [L3T−1] is the total sediment flux leaving the cell and Qt [L3T−1] is the sum of the transport capacities in all lower directions. The flux qsj [L2T−1] is the sediment flux leaving the cell in direction j, wf [LT−1] is the fall velocity of clasts in still water, Ksa is a bedrock detachment coefficient, ρw is the density of water and a and b are exponents. The incision rate Ij is the maximum for a shear stress value that falls between the critical shear stress for clast entrainment and a value for the passage from saltation (efficient for abrasion by impacts) to wash transport.
 Changes in elevation Δz are approximated by a finite volume approach [Tucker et al., 2001]
where U [LT−1] is the rock uplift rate, Qsin [L3T−1] is the total sediment volume flux entering the cell, and the sum applies to all lower neighboring cells j. Also, Δt is the time increment, and the flow width Wj is determined by equation (5) in the case of water-driven transport or by the length of the side of the inset octagon within a cell (( − 1)Δx) in the case of small transport distance processes (equation (3)). Sediment outflux per unit width qsj toward a lower cell j could be the result of sediment or bedrock erosion, or both. We describe this process on one cell in the following.
 The transport and detachment rates provided by equations (3), (4), and (6) or (7) are considered to be maximum possible values. First a volume eroded by nonlinear diffusion (equation (3)) is calculated. If no sediment remains, nonlinear diffusion applies to bedrock. Then, the stock of sediment available for water-driven transport is calculated by taking the sum of sediment already in the cell and the total incoming sediment. This stock is compared to the sum of transport capacities calculated for each flow pathway leaving the cell [e.g., Thomas et al., 2007]. If the total transport capacity is higher than the amount of available stock, the bedrock is eroded (equation (6)), but the detached volume (the sum of detached volumes heading downstream) cannot exceed the remaining capacity. Thus, at every point in time, erosion can be limited by bedrock detachment (“detachment-limited” erosion [Howard and Kerby, 1983]) or by transport capacity (“transport-limited” erosion [Willgoose et al., 1991a]) depending on the model's parameters and on the evolution of the topography at every point.
 The finite volume approach adopted in our model (equation (8)) results in an underestimation of the river bed elevation variation. Indeed, the eroded or deposited volume is spread out over the entire cell area, which is larger than the river bed area [Tucker and Slingerland, 1994; Carretier and Lucazeau, 2005; Loget et al., 2006]. Thus, with equation (8), the cell elevation is defined as the mean elevation of the cell. Alternatively, we could have calculated Δz by dividing the volume of sediment by the flow width, assuming that the cell elevation corresponds to the river bed. However, this would result in overestimating the volume gained or lost by the topography, because the river bed variation would be applied to the entire cell, even though part of the cell contains a hillslope. Consequently, the accumulated eroded volume measured from the topography would be greater than the volume measured by integrating the outflux of sediment at the mountain river outlets. Neither of these two approaches is perfect, and there has been little discussion of the differences between them.
Some reference experiments are repeated in order to facilitate the comparison with other ones. The code “ic” is for initially connected surface. These experiments start off from a dynamical equilibrium topography and a doubled uplift rate. Other values of the initial slope are measured in degrees. The unit Kbr is m s−1 Pa−a. Kbr = means that sediments are always fully available (transport-limited). The unit Kall is m2 s−1 Pa−p. Kall = means that the transport capacity is infinite, so the volume detached from bedrock is only limited by bedrock resistance (detachment-limited). The unit τc is Pa. The letters for each type of model correspond to E, exponential; SL, straight line; L, linear; S, sigmoid. E5 appears twice with different m, n, and p values, which are strictly consistent and help for comparing with other experiments.
 In one such reference experiment (E2), we must first justify the parameter values. This experiment applies to a 50 × 60 cell grid (cell width of 250 m), corresponding to an initial horizontal surface with Gaussian noise (σ = 0.5 m). Alluvial transport and bedrock detachment laws (equations (4) and (6)) are power functions of the excess shear stress and assume steady, uniform flow in a wide channel. Using the Manning flow resistance equation, m = α = 0.6 and n = β = 0.7 in equations (4) and (6) [see, e.g., Tucker, 2004].
 The excess shear stress exponent of the transport capacity p = 1.5 and the transport coefficient Kall = 1.5 10−5 m2 s−1 Pa−1.5 (equation (4)) correspond to the Meyer-Peter and Muller  formula. The value of τc is 5 Pa, which corresponds to grain size of about 1 cm in diameter.
 The flow width parameter kw = 12 m−1.5 s0.5 results in a river width of about 20 m at outlet. Finally, the equation for hillslope processes uses classical values for the diffusion coefficient κ = 10−2 m2 a−1 [e.g., Martin and Church 1997] and the slope stability threshold Sc = 0.6 (=tan 30°) for sediment and Sc = 1.7 (=tan 60°) for bedrock.
 Our analysis focuses on the effects of the following factors (Table 2): (1) the transport threshold τc. The tested values (<30 Pa) correspond to grain sizes ranging between fine sand and 5 cm in diameter, (2) the system size, ranging from 15 to 37.5 km in length, and (3) the ratio between the incision and transport coefficients Kbr and Kall, which partially controls the model's tendency to be either transport or detachment-limited during the transient response [Howard, 1980; Whipple and Tucker, 2002; Cowie et al., 2006]. The values tested lead to a realistic mountain relief (between 200 m/15 km and 1500 m/15 km for drainage areas of about 100 km2). Some experiments are entirely transport-limited and others are entirely detachment-limited, (4) the nonlinearity of erosion-transport laws in slope and discharge, with the parameters m, n, α, β, p and a in equations (4) and (6). The values tested for α, m, β, n, p and a are all within the expected range [0.5–2] for many bedrock erosion and transport laws [Whipple et al., 1998; Yalin, 1972] and (5) the initial topography. The initial surfaces tested are either horizontal, gently inclined planes, or a topography equilibrium of another experiment from which the uplift rate is doubled.
 Experiment E41 uses the same parameters for a = −0.5, b = 1.5 (equation (7)) from Sklar and Dietrich's  law. The fall velocity in water, wf = 0.3 m s−1, corresponds to sediment grain size of several centimeters wide, and consistent with τc = 30Pa. The detachment coefficient Ksa is larger (10−4 m−1) than the value of Sklar and Dietrich  to avoid unrealisticly large slopes for small drainage areas [see Gasparini et al., 2007].
 All simulations use periodic boundary conditions linking the western and eastern sides of the model grid, a wall boundary (no output flux) on the north side and free output flux on the south side. The same uplift rate U (1 mm a−1) and precipitation rate P [LT−1] (=1 m a−1 in all experiments) are applied uniformly to the entire domain. Starting from a specific initial surface, the model runs until a dynamic equilibrium is reached. During the transient stage, we record the evolution of the topography and erosion rate of the whole model and of selected rivers, and compare their evolution with the erosion laws under evaluation.
 In the following analysis, we use the terms “complete connectivity” or “initially connected surface” [e.g., Davy and Crave, 2000; Lague et al., 2003]. For an initially connected surface, we designate a surface in which all cells are connected to the base level through a drainage network, which remains unchanged after the onset of uplift. Note that for a planar dipping surface, all pixels are connected to the base level, but the growth of the catchment growth then modifies the drainage network. Consequently, such a surface is not called here an initially connected surface.
4.1. Large-Scale Erosion Rates
 Experiments typically show two stages of the topographical adjustment, as illustrated by Figure 1 for three experiments (E30, E27, E2): the first stage takes place while the drainage system is developing, and the second includes continuing evolution of the topography toward a dynamic equilibrium. The first stage lasts for an amount of time Tco (connectivity time), and the total response time to achieve the dynamic equilibrium from the onset of uplift is called T. Tco is estimated by the time at which there are no close depressions any more in experiments starting from a horizontal surface, and by the time at which channel heads reach the northern side in experiments starting from a dipping surface. T is estimated by the time at which the average spatial erosion rate ε(t) = 0.99U or ε(t) = 1.01U, depending on the experiment type.
 We first investigate the average spatial erosion rate ε(t) of the uplifting block, defined as the total sediment output flux on the southern side divided by the area of the uplifted domain. Figure 2 and Table 2 sum up the results. Four typical previously observed ε(t) curves can be identified, some having emerged in numerical models and others in physical models: The first is known as the “linear”-type because it has the form of a linear response function ε(t) = U(1 −exp(−t/T)) [e.g., Kooi and Beaumont 1996]. The second characteristic ε(t) curve is the “sigmoid”-type. This sigmoid curve has been observed, for example, in both physical experiments [Hasbargen and Paola, 2000] and numerical simulations [Carretier and Lucazeau, 2005]. The third ε(t) curve is the “straight-line”-type. Finally, the fourth characteristic model is the“exponential”-type with a concave up increase of ε(t) and a transient maximum that is greater than the uplift rate [e.g., Lague, 2001]. As we examine in detail in the following, these model types depend strongly on Tco.
 Linear-type curves approximately obey an equation of the type ∂ε/∂t = (U − ε)/T, which is the equation of a linear system with input U and output ε [Kooi and Beaumont, 1996]. Linear-type models have a negligible connectivity time Tco compared to T. This is pertinent for some of the experiments that start from an inclined plane (E30), all the experiments starting from a previous dynamic equilibrium that double the uplift rate, no matter what the value of τc is (E28, E31, E35, E40), and one transport-limited experiment that uses a threshold-free transport law linear in discharge and slope (E5). In the latter, drainage connections form between depressions on the plateau, which is similar to experiment E27 as shown in Figure 1b. This creates depressions with larger area, and their capture helps the drainage network develop quickly. Experiments that are entirely transport-limited and those including detachment-limited erosion can also be linear (e.g., E30 and E28). For experiments in which erosion can be detachment-limited, increasing the initial regional slope does not necessary imply a negligible Tco (e.g., E4, E17, E32, E33, E34). From a certain regional slope, Tco even remains constant (E33, E34 and E37, E38), which suggests that detachment-limited erosion imposes a significant connectivity time to establish the drainage network. Note that no closed depression exists in any of the experiments that starts from a dipping surface, because the regional slope dominates the gaussian noise added to initial elevations.
 The conditions required for a linear-type model are consistent with Kooi and Beaumont's  conclusions. The linear behavior of the whole system essentially requires that the drainage network does not change after the onset or acceleration of uplift. This is true for experiments starting from a previous dynamic equilibrium. For other linear-type experiments, the drainage network is established very quickly (see Figure 1a). Strictly speaking, linear behavior should be valid for erosion/transport laws linear in slope only [Kooi and Beaumont, 1996; Lague, 2001]. However, our experiments show that quasi-linear behavior can emerge from laws that incorporate a positive τc and that are thus nonlinear in slope (E28, E30, E35, E40). This is the case for some experiments starting from a dipping surface (E30) or from an initially connected surface (E28, E35, E40). For these experiments, the nonlinearity of erosion/transport laws is not the main factor controlling the shape of the ε(t) curve.
 Straight-line-type models combine two characteristics: (1) a significant Tco and (2) very low river concavities. In these models (E1, E15, E16), Tco is large because the drainage network development rate is limited by bedrock detachment at channel heads. However, the erosion tends to be transport-limited at the dynamic equilibrium, so that the parameters of the transport capacity determine the river concavity. The river concavity at dynamic equilibrium is also the exponent θ in the slope-area relationship S ∝ A−θ, obtained by equating the tectonic flux and the sediment flux (equation (A5)) [e.g., Flint, 1974; Tarboton et al., 1989; Willgoose et al., 1991b]. The parameters of the transport capacity used in straight-line-type experiments lead to a very low concavity θ = 0.05 that implies a very smooth topography and straight rivers [Kirkby, 1971; Tucker and Whipple, 2002]. A concavity of 0 is generally not observed in real or experimental river profiles [Whipple and Tucker, 2002; Lague et al., 2003] (except for those rivers that receive water only at the highest elevations). Large concavity values can be associated with significant τc [Tucker, 2004]. There is a growing number of arguments pointing to the significant effect that the transport threshold τc has on the erosion dynamic of mountain belts [Parker, 1978; Snyder et al., 2003; Talling, 2000; Lague et al., 2003]. This arguments suggest that the straight-line-type model is unrealistic. We point out that parameters of experiment E5 lead to θ = 0 but E5 has a linear-type behavior. In this experiment, transport-limited erosion is imposed, contrary to the straight-line-type experiments that incorporate detachment-limited erosion. Thus, the drainage network develops very quickly in E5, which leads to a linear-type model.
 Sigmoid-type models are characterized by a significant Tco value that corresponds to the time before the inflexion point of the sigmoid curves (Figure 2). This long Tco implies a wave-like behavior during the connectivity phase, and a more diffusive behavior after (Figure 1b). The growth of the drainage network is responsible for the nonlinearity of the whole system's response as measured by the mean erosion rate [see also Kooi and Beaumont, 1996]. These models include detachment-limited experiments (E36, E37, E38), experiments using a significant τc > 1 Pa (E17, E18, E29, E32, E33, E34), and transport-limited experiments using a threshold-free transport capacity that is more than linear in water discharge (E10, E27). The common point in these experiments, as we explain in detail in section 5.2, is that the sediment flux (the transport capacity in the transport-limited experiments or the upstream integral of detached sediment in detachment-limited experiments) increases more than linearly with water discharge. Sigmoid-type models appear to be intermediate, falling between the linear and exponential types.
 Exponential-type models correspond to models with a much more significant period of connectivity (typically Tco > 0.5 T see Figure 2), and for which the mean erosion rate passes through a maximum that is higher than the uplift rate. During the drainage network growth, ε(t) can be approximated by a function of the type (et − 1) or equivalently by a power law function of t with an exponent >1. Exponential-type models result from the same kind of erosion and transport laws as the sigmoid-type models. In the case of transport-limited, threshold-free models, the transient excess of the mean erosion rate above the uplift rate is almost imperceptible (Figure 2a), even when the length of the system length is more than doubled (E11, E12). This difference is explained by (1) the intrinsically diffusive character of transport-limited erosion that limits the rate of sediment exportation in catchments and thus prevents the mean erosion rate from significantly exceeding the uplift rate [see also Lague, 2001] and (2) the fact that endoreic zones get connected on the plateau, which favors the drainage network growth rate (Figure 1b). This second point is true if the value of the slope exponent n of the transport capacity law is not too much greater than 1. With higher values of n, the transport capacity decreases dramatically for very low slopes (smaller than 1) on the plateau, which prevents the development of the drainage links.
 In summary, quasi-linear models occur if Tco = 0, sigmoid-type models occur if Tco < 0.5T, and the exponential-type models occur if Tco > 0.5T. Figure 2b sums up the relationships between model type, associated conditions and the Tco/T ratio. There is not further discussion of the straight-line model, as it appears to be unrealistic.
4.2. Relationships Between Large-Scale and Local Erosion Rates and River Profile Evolution
 If the local incision rate in a river exceeds the uplift rate, does this imply that the mean erosion rate is of an exponential nature? Is there a relationship between the type of mean erosion rate evolution and the shape of transient river profiles? To answer these questions, we extracted the profiles of major rivers over time from each experiment, and plotted the evolution of the local incision rate for two points located on the first and second thirds of the river (P1 and P2). The results are summarized in Figure 3, which displays the large-scale mean erosion rate ε, the large-scale mean elevation (z), the profile adjustment of a selected river and the evolution of the incision rates (I) at these two selected points. The three cases shown in Figure 3 correspond to the experiments illustrated in Figure 1 (E30 for the linear-type, E27 for the sigmoid-type, and E2 for the exponential-type). The following differences can be observed:
 1. None of the models shows a concave up increase in the mean elevation curve during the development of the drainage network. The mean elevation of the exponential-type reaches a maximum, but this occurs before the maximum of the mean erosion rate.
 2. The river profiles evolve differently for each type of model. In the linear model the profile evolves by increasing the slopes. In the sigmoid-type model, the river profile evolves by first decreasing and then increasing the slopes. In the exponential-type model, the river profile evolves by decreasing the slopes.
 3. In the case of the linear-type model, the local incision rates of the two points never exceed the rock uplift rate, whereas they can reach much greater values than the rock uplift rate in the two other cases. However, the time at which this local maximum occurs corresponds to the passing of a retreating erosion wave [Stark and Stark, 2001], and thus is not related to the time at which the maximum mean erosion rate occurs (in the exception of the points located at the catchment head). Moreover, the farther the point is located within the catchment, the more pronounced is the spike of the incision rate. This reflects the fact that disequilibrium increases toward the channel head.
 These relationships between the evolution of the mean erosion rate, local erosion rate and transient river profiles were observed in all our experiments. In the case of the linear-type model, the river profile can show a wave-like behavior during its evolution, which is different from the example displayed in Figure 3a. This occurs for detachment-limited models that start from a previous equilibrium and that double the uplift rate. However, even in this case, the relationships described above remain true. So it seems that, once again, the ratio between Tco and T determines the style of the topographic adjustment for all scales.
 The difference between the shapes of the mean erosion rate and the mean-elevation curves, as well as the shift between the maximum mean erosion rate and the maximum mean elevation in the exponential-type model, can be explained rather simply. Indeed, the mean elevation z is
Thus the maximum of z occurs when dz/dt = U − ε(t) = 0, which is when the erosion rate equals the uplift rate for the first time (Figure 3).
4.3. Implication for the Field Interpretation of Terraces
 The three styles of river adjustment that have been identified may have implications for interpreting field data (Figure 4). If we consider that the successive river profiles correspond to terraces abandoned by climatic or sediment-driven river width variations, then the downstream or upstream convergence of terrace long profiles can be strongly influenced by the length of the connectivity phase. In linear-type models, terraces must converge upstream, while in exponential-type models terraces converge downstream. The sigmoid-type models are capable of generating terraces that converge both downstream for the oldest terraces and upstream for the youngest ones (Figure 4b). However, these predictions are made for temporally and spatially constant uplift rate and precipitation, whereas real systems can be much more complex [Molnar et al., 1994; Merritts et al., 1994; Pazzaglia and Brandon, 2001; Poisson and Avouac, 2004].
 We tested the identified relationships on a well-studied field example located on the Ih Bogd Mountain, in Mongolia [Ritz et al., 1995; Bayasgalan et al., 1999; Carretier et al., 2002; Ritz et al., 2003; Vassallo et al., 2005, 2007b, 2007a; Jolivet et al., 2007]. The Ih Bogd Mountain is a 50 × 25 km2 transpressional ridge associated with a major 260 km long strike-slip fault system. It is bounded by reverse fault segments. Like many other mountains in this area, Ih Bogd has conserved a large part of the undissected, preuplift surface remnant in its core [Jolivet et al., 2007] (Figure 5), showing that this massif is still in the drainage growth phase (t < Tco), which is particularly long considering that uplift began about 5 Ma ago [Vassallo et al., 2007a]. This surface dips very gently (<1°) toward the Bitut Valley, without any trace of channelized flow. The soil has periglacial characteristics, with a roughness of several tens of centimeters, and small (10 m x 10 m) and shallow depressions (<1 m) with desiccation traces. The evaporation and infiltration rates are unknown but possibly significant given the very low regional slope of the plateau. The drainage network is clearly retreating into the plateau, with channel heads showing a sharp slope break [Vassallo et al., 2007a]. Topographic surveys and cosmogenic nuclide dating from the Bitut river terraces allowed Vassallo et al. [2007a] to quantify incision rates between terraces. Dating suggests that terraces formed every 100 ka, so that the incision rate should integrate climatic variations, as assumed in our theoretical study. Independent estimates of uplift rates on bounding faults allowed [Vassallo et al., 2007a] to compare the Pleistocene uplift rate (ca 0.1 mm a−1) with the Pleistocene incision rates. Uplift rates for both sides of the massif are similar, and the core surface is almost horizontal, which suggests that the uplift rate is spatially constant. Incision rates are 3 to 6 times larger than the uplift rate (Figure 5), which Vassallo et al. [2007b] interpreted as being mainly the result of drainage growth, and also of the interaction with piedmont and cyclic climatic variations [Carretier and Lucazeau, 2005]. Figure 5 shows that Bitut terraces converge downstream. This observation, along with the fact that the local incision rates are greater than the rock uplift rate and the existing remnant of the preuplift surface are all consistent with an exponential-type model. All this supports Vassallo et al.'s  conclusions that in this case, most of the topographic adjustment to rock uplift for different scales is controlled by the ongoing growth of the drainage system. This also suggests that the long timescale erosion is either detachment-limited or transport-limited with a positive transport threshold, which seems to be consistent with the coarse sediment cover and exposed bedrock in some parts. Catchment-scale erosion rates are not yet available so it is not possible to prove the full consistency between this example and the proposed theory.
5.1. Analogy With Biological Population Growth and Percolation
 Drainage network growth has been shown to be analogous to invasion-percolation models on the basis of similar fractal dimensions of the drainage network [Stark, 1991; Yakovlev et al., 2005]. Moreover, Stark  studied the relative effects of three processes and elements involved in the retreat of a drainage front: weathering and weakening of the substratum, seepage erosion and initial rock strength. To take these components into account, he used a phenomenological modeling approach based on network growth theories including invasion-percolation, diffusion-limited aggregation (DLA) and Eden growth (parallel retreat). Varying the strength of each process, he showed that the drainage front line is more or less irregular with fractal dimensions depending on the processes involved. Similar front shapes have been observed in our experiments, though varying in time and between different experiments. Such shapes include uniform retreat (Eden growth) and localized channel head invasion, although seepage erosion is not taken into account (Figure 1). In our simulation, erosion retreat proceeds either by reducing the slope to a critical slope (equation (3)), which favors Eden growth, or through fluvial erosion, depending on the transport threshold τc. However, bedrock collapse is not the dominant process, as model slopes are generally much smaller than Sc. In any case, just as in Stark's  model, network growth depends on the initial cell property (initial slope and associated drainage area) and processes that weaken the rock resistance to erosion, and which can evolve in time. The latter corresponds to the formation of drainage links on the plateau or the capture of depressions that suddenly increase the erosive power of cells located just downstream the captured depression. A visual inspection of the different shapes of the erosion front did not show a clear correlation between a particular shape and model type, in particular because the mode of channel retreat varies with time. Nevertheless, Eden growth seems to be more pronounced in sigmoid and exponential-type models, as shown in Figure 1. We did not carry out a fractal analysis due to the coarse resolution of the model grid.
 Here we make another comparison. In analyzing the sigmoid-type models, it can be noted that the ε(t) curve follows a Weibull curve, defined as (Figure 6):
where γ is the inverse of a characteristic time and μ is a positive constant that lies between 1 and 2.5 in our experiments. If μ = 1, ε(t) is a linear-type model. The connectivity time Tco corresponds to the maximum slope of the Weibull curve, so that
 It is interesting to note that such sigmoid curves (Weibull, Logistic, etc.) are known in biology to represent population growth processes in a context of competition for food in a limited space, which are often modeled as percolation models. The eroded volume at each moment in time might be analogous to population growth. If this analogy is correct, then the connectivity process is limited by the potential to erode (the food), as this potential is limited by bedrock detachment or an entrainment threshold.
 The exponential-type models deviates from the Weibull curve. This may be due to a fundamental difference with the population analogy: in the case of an actively uplifting mountain range, the erosion rate seems to be able to make “reserves”, which is to say that some material provided by rock uplift is stocked for later. This can lead the mountain to an unstable state in which its erosion rate and mean elevation are greater than the equilibrium state. This is not observed in population growth studies because populations do not usually make food reserves over long periods of time. In the case of a mountain, the potential erosion can be stocked up during the connectivity stage because slopes are abnormally large as a consequence of the small growth rate of the drainage area. When the drainage area reaches its maximum, slopes relax and the mean erosion rate decreases.
5.2. What Does Limit the Drainage Network Growth?
 In order to study the effect of the parameters on connectivity time, we first compare the ratio between the connectivity time and the response time (Tco/T), or the connectivity time alone with nondimensional numbers (see Appendix A and Figure 7). Numbers Nd and Nt measure the contribution of τc to an equilibrium slope. They are, in other words, the ratio between the capacity to erode and the resistance (τc) for a specified drainage area at dynamic equilibrium, in the cases of detachment and transport-limited models, respectively [Tucker, 2004]. The larger these numbers, the smaller the threshold effect. Number Ndt determines if the system is detachment-limited (Ndt > 1) or transport-limited at dynamic equilibrium for a given drainage area [Whipple and Tucker, 2002]. It helps evaluate the potential effect of the dominant erosion-limiting process on connectivity time. Figures 7a and 7b show that Tco/T tends to decrease with Nd and Nt, suggesting that τc increases the connectivity time significantly [see also Lague, 2001], which also appears when plotting Tco against τc (Figure 7d). The Tco/T ratio does not demonstrate a clear relationship with Ndt. These plots show a significant variability, suggesting that no one of the evaluated processes can explain the Tco/T ratio alone. Moreover, these numbers measure a balance of processes during the steady state, which may not hold during the transient stage. For example, even if transport-limited conditions are predicted at dynamic equilibrium, detachment-limited conditions can prevail in a large portion of catchments during the transient response [see also Tucker and Whipple, 2002].
Tco increases linearly with the size L of the system for experiments with τc = 0 (E8, E9, E4) and it increases as L0.8 for experiments with τc ≫ 0 (E10, E11, E12). A more or less linear increase has also been observed in the case of threshold-free, transport-limited erosion models of an initial plateau [Davy and Crave, 2000], and in sandbox erosion experiments of an uplifting block [Rohais, 2007]. This linear increase highlights constant drainage growth rate [Davy and Crave, 2000]. To the contrary, Lague et al.  observed a rate that decreased over time, but this difference could have been caused by a difference in boundary conditions: the four sides of the block are open in Lague et al.'s  experiments, implying that the connectivity of the overall system may be limited by the competing growth of catchments that run in perpendicular directions. In the other cases, constant drainage network growth rates have been obtained for different erosion laws and initial random surfaces. This suggests that other processes control the stability of the drainage growth rate.
 In some cases, Kbr influences Tco, and in other cases it does not. For example, Tco decreases with Kbr in E4, E13, and E14 (large τc) and in E1 and E15 (small τc), but it is unchanged in E17 and E18 (positive regional slope and large threshold). In this last case, the initial dipping surface seems to control Tco. In the case of a small threshold case (E1 and E16), decreasing the transport capacity coefficient Kall strongly increases Tco, which confirms the intuitive notion that the rate at which sediments are evacuated influences the rate of drainage network creation. This rate depends also on the excess shear stress exponents a and p. Increasing a or p will increase the erosion/transport capacity when τ ≫ τc, and thus enhance deep river incision or facilitate sediment exportation. This tends to decrease the connectivity time Tco, as illustrated by the comparison between experiments E2–E21 (p increase) and E20–E2 (a increase).
 The nonlinearity of erosion/transport law in discharge and slope may provide a better framework to understand how the erosion rates and topography evolutions observed in our study are controlled. In the sigmoid and exponential-type models, long Tco are associated with a sharp retreating front with concave-up river profiles at the channel head (Figure 3). Such a feature tends to arise under any or all the following conditions: (1) in a detachment-limited model using a stream power law with a slope exponent larger than 1 (I ∝ (Q/W)αSβ, β ≫ 1) [Tucker and Whipple, 2002], (2) in a transport-limited model using a transport law with discharge and slope exponents much larger than 1 (qt ∝ (Q/W)mSn, m ≫ 1 and n ≫ 1), and (3) in models using erosion/transport laws that incorporate a positive threshold [Tucker, 2004]. As discussed in section 4.1, a slope exponent n greater than 1 limits drainage connections in the plateau and induces long Tco. In the case of a detachment-limited model, β > 1 creates a slope break at the plateau which prevents endoreic zones from being captured. Moreover, Tco significantly increases with the discharge exponents m in a transport-limited model (E5, E27) and α in a detachment-limited model (E24, E25, E26). The common point to both cases is that the volume sediment flux Qs grows more than linearly with catchment area A (or water discharge Q) [Smith and Bretherton, 1972; Tucker, 2004], thus allowing for a deep incision in growing catchments. If this incision is large enough, hillslope gradients can increase to the point that the mean erosion rate exceeds the uplift rate [see also Lague, 2001].
 In the detachment-limited and threshold-free models (I ∝ A0.5), the rapid downstream increase of sediment flux is true by definition, because the transport capacity is implicitly assumed to be always equal or larger than the upstream integral of eroded sediment. At the river outlet, the sediment outflux Qs is the integral of the erosion rate I over the drainage area. Therefore, Qs is a function of A raised to a power >1. Because A increases linearly during the drainage network development, Qs(t) (or ε(t)) at the river outlet is also a power function of t raised to a power >1. This is what is observed for the sigmoid-type and exponential-type models during the drainage network growth phase. Indeed, ε(t) can be approximated by a function of type (et − 1) or equivalently by a function of type tr with r > 1. In other words, the shape of the curve Qs(A) mimics the shape of the curve ε(t). The same relationship applies to the transport-limited and threshold-free case if m ≫ 1.
 In transport-limited modes with a significant transport threshold, [Tucker, 2004] showed that the transport capacity increases more than linearly with the drainage area. So again, the transport law curve Qs = f(A) mimics the ε(t) curve during drainage growth.
 In addition, Davy and Crave  found a similarity between transport-limited models in which m≥2 and detachment-limited models (laws linear in slope and with no threshold in both cases). In both cases, the characteristic time necessary to erode an initial plateau does not depend on the size of the plateau, though this is not true if m < 2. Also in both cases, the sediment flux is much more than linear in the drainage area, which allows for sediment to be exported over a long distance.
 This analysis applies to the straight-line models too. Indeed, the linear increase of the mean erosion rate corresponds to the linear scaling between the transport capacity and the water discharge.
 Analyzing the relationship between local erosion/transport laws and sediment outflux during drainage growth may be one way of evaluating the form that these laws take in physical models or in nature using sedimentological data. For example, [Hasbargen and Paola, 2000] found a more-than-linear increase of sediment outflux during the transient stage of their uplifting sandbox experiments, which is similar to the sigmoid or exponential-type models. This suggests a strongly nonlinear relationship between the erosion/transport law and water discharge. Moreover, such an increase has also been observed in reconstructed sediment outflux at Tibetan outlets [Metivier et al., 1999; Clift, 2006]. This could correspond to our exponential-type models, although it could also correspond to temporal variations in uplift rate or climate.
 It is important to point out that taking the dependence of river width on slope into account, as proposed by Finnegan et al.  and Whittaker et al. [2007b], is implicit in some of our experiments. In fact, the incorporation of this dependence essentially leads to the same equations (4) and (6), with a 20% to 40% larger slope exponent [Attal et al., 2008]. For example, the threshold-free and detachment-limited experiments E25 and E26 use larger slope exponents than E24. A larger slope exponent tends to increase the connectivity time because, as we previously discussed, a large slope exponent prevents drainage links from forming at the channel head. This effect seems to dominate the competing acceleration of incision in gorges beneath the channel head. In the case of a normal fault acceleration, Attal et al.  showed that a decrease in channel width with an increasing slope accelerates the topographic response. This appears to be different for an initially random surface combined with a symmetrical uplift. The effect exercised by the slope exponent can also be evaluated in the case of a significant τc (E22 and E23). Similar connectivity times are obtained in both cases, probably because τc is the key parameter controlling the nonlinear relationship between the erosion law and the slope [Tucker, 2004]. In addition, a significant bedrock river width adjustment time can also influence the connectivity and the entire system response time [Stark, 2006]. This must be evaluated further.
Gasparini et al. , Gasparini et al. , and Sklar et al.  showed that the use of bedrock incision models that incorporate the tool and cover effects [Sklar and Dietrich, 2004] significantly modifies the river profile adjustment to an increase in uplift. This can potentially yield different behaviors than those observed in our study. However, if the growth of the drainage system controls most of the system response, the behavior of sophisticated bedrock incision models might fall under the model classification that we have proposed. This is illustrated by experiment E41, which uses [Sklar and Dietrich's,  law and leads to an exponential-type model. The rationale could be that sediment-flux-dependent bedrock incision models converge on a transport-limited model for large drainage areas [Gasparini et al., 2007]. This would mean that the nonlinearity of the transport capacity in discharge and slope may control the transient evolution of the mean erosion rate during drainage growth. Note that Gasparini et al.  also observed that the transient mean erosion rate exceeded the uplift rate. Nevertheless, further analysis is required to evaluate the potential effect that erosion laws incorporating tool and cover effect may have on connectivity time and the associated river profile adjustment.
 Finally, our hydrological model assumes that water disappears in lakes. Allowing water to fill lakes would probably decrease the drainage connectivity time, favoring linear-type rather than sigmoid or exponential-type models. However, experiments that started from a dipping surface did not contain any lakes. Significant connectivity times were observed in these cases, demonstrating that the connection of endoreic zones is not the limiting process in this case. To the contrary, the hydrological behavior of a horizontal plateau could represent a fundamental (and often neglected) controlling factor of the erosion dynamic [e.g., Hovius et al., 1998].
5.3. Relation With the “Scarp Retreat” and “Down-Wearing” Scarp Evolution Models
 Different styles of plateau erosion have been classified in two end-member cases: the “scarp retreat” or “back-wearing,” and the “down-wearing” models [Kooi and Beaumont, 1994; van der Beek et al., 2002; Tucker and Whipple, 2002]. The scarp retreat model corresponds to the horizontal removal of plateau material through the propagation of an erosion wave, whereas the down-wearing model corresponds mostly to the vertical erosion of the plateau. Tucker and Whipple  showed that these models could be strongly associated with the slope exponent in erosion law. In the case of a detachment-limited situation, if the erosion rate scales to slope with an exponent α > 1 then scarp retreat is favored, whereas if α < 1, down-wearing dominates. Moreover, van der Beek et al.  showed that these two styles are also strongly influenced by the initial slope of the plateau. The existence of an inland drainage divide favors down-wearing because it allows for the rapid development of the drainage network on a surface dipping toward the base level, and erosion can occur everywhere on the plateau. To the contrary, scarp retreat occurs if the initial topography dips away from the scarp. In other words, the style of the plateau erosion is influenced by the connectivity time, which is consistent with our experiments: a dipping initial surface led to the down-wearing style and linear-type model, whereas a horizontal initial plateau implied a strongly horizontal component of erosion, associated with the sigmoid and exponential-type models (exponent, Figure 1). This suggests that the scaling of the erosion law to the slope does not necessarily control the erosion response to uplift if other factors, such as infiltration or the erosion threshold, control the connectivity. In particular, the style of plateau erosion may not be characteristic of a particular slope exponent of the erosion law if the erosion threshold is large. Indeed, a large threshold will mainly enhance the nonlinearity between the erosion rate and slope [Snyder et al., 2003; Lague et al., 2003; Tucker, 2004]. For example, experiments E22 and E23 (detachment-limited) or E2 and E19 (detachment and transport-limited) used detachment-limited erosion laws with erosion thresholds, but differ by their slope exponent. The connectivity time is the same in both pairs of experiments, showing that it is more strongly controlled by the erosion threshold than by the slope exponents.
 Our study points out that the growth of the drainage network could exert a primary control over the landscape response time to uplift [see also Kooi and Beaumont, 1996; Lague, 2001; van der Beek et al., 2002; Lague et al., 2003]. From numerical modelings of uplifting block erosion, we have identified three main realistic model types for the evolution of the mean erosion rate. These are the linear-type, the sigmoid-type, and the exponential-type models. Linear-type models are obtained for initially connected surfaces. Sigmoid-type (Tco < 0.5 T) and exponential-type (Tco > 0.5 T) models correspond to evolutions with long connectivity times. The mean erosion rate can exceed the uplift rate in exponential-type models.
 Moreover, linear-type models yield river terraces that converge upstream, whereas exponential-type models yield terraces that converge downstream. Thus, by documenting the long-term sediment fluxes out of mountains and river terrace geometry, we should be able to better constrain the processes limiting erosion.
 In the case of the linear-type model, the local incision rate in the river never exceeds the uplift rate, in contrast to the other two cases. Thus, the large-scale erosion rate and the local incision rate could be lower or greater than the uplift rate at the same time in an uplifting block, which could explain differences in the measurements of erosion rates at different spatial scales in a catchment.
 Whatever the controls, connectivity time is still not totally understood, especially concerning uplift rate, sediment-flux-dependent incision and hydrology. With our model setting, long connectivity time occurs when the transport capacity is more than linear in drainage area and slope, particularly in the case of detachment-limited models, and in the transport-limited models using a positive transport threshold. To further understand what controls the connectivity time from a quantitative point of view could provide us with useful information on the erosion laws both in the real world and in physical experiments.
Appendix A:: Nondimensionalization
 From equations (8), (4) and (6) we can write relationships between all variables at dynamic equilibrium (Δz/Δt = 0) for two end-members models, namely, the detachment-limited model
and the transport-limited model (all is sediment):
where A [L2] is the drainage area. Assuming no infiltration and homogeneous rainfall Q = PA, in both cases equilibrium slopes can be deduced as a function of A.
 In the detachment-limited model,
where Sd is the detachment-limited equilibrium slope. In parentheses both terms of the sum represent what Tucker  named the capacity and competence terms, respectively. Dividing the first term by the second leads to a nondimensional number Nd, the detachment/competence number that measures the ratio between the capacity to erode and the resistance of the bedrock [Tucker, 2004]:
 Similarly, from equation (A2) we obtain the following for the transport-limited model:
where St is the equilibrium slope, and
where Nt is the capacity/competence number for transport.
 Dividing Sd by St we obtain another nondimensional number Ndt the value of which determines if the erosion is transport (Ndt < 1) or detachment limited (Ndt > 1) at equilibrium as a function of the drainage area A [Whipple and Tucker, 2002]:
 This study was funded by the IRD, the French national research agency ANR (project “ANDES” ANR-06-JCJC-0100), and the Chilean Anillo (project ACT-18). We thank M. Attal, C. Stark, P. van der Beek, and the Associate Editor G. Tucker for outstanding reviews which improved this paper. S. Carretier and E. Pepin thank the Department of Geology of the University of Chile for its warm welcome.