Surveyed outcrops of the Middle Jurassic Entrada Sandstone at Ghost Ranch, New Mexico, show the unusual occurrence of preserved aeolian dune palaeotopography buried beneath subaqueous strata. The preserved dune remnants have relief up to 35 m, trend NNW, and show internal scalloped cross-strata dipping to the WSW, with small sets occurring as both topsets and bottomsets. Outcrop data are best satisfied in computer models by 50 m high, sinuous bedforms that migrated to the WSW, while the sinuosity migrated alongcrest to the NNW. Superimposed small dunes occurred upon the stoss slope, and at the basal lee of the main bedform where they migrated alongslope to the NNW. Remnant dune palaeotopography is buried by onlapping, subaqueous, largely structureless sandstones believed to be derived by mass wasting of the upper portions of the dunes and deposited as sediment-gravity flows that infilled between the dunes. Preservation of dune palaeotopography beneath mass-flow deposits, with no evidence for gradually rising water, argues that flooding of the Entrada dune field was geologically instantaneous. The thickness and lithology of the overlying Todilto Formation conform to slight remnant palaeotopography on the Entrada surface. The Todilto is a laminated limestone and thinnest over remnant dune crestal areas, but thickens and increases in gypsum content downslope until it abruptly yields to a gypsum mound positioned over a remnant interdune hollow. The Todilto laminations are interpreted as seasonal varves deposited below wave base in a density-stratified water body. The flooding event that gave rise to the controversial Todilto water body occurred during Entrada time, with Todilto deposition occurring within an already substantial water body.