We thank E. Schweitzer, M. Schibler, and Y. Song for advice on immunostaining and B. Sjostrand for technical help on electron microscopy. We also thank J. Tian for advice on the generation of GABARAP antibodies. This study was supported by grants NS35985, NS28772 (to R.W.O.), and training grant GM07185 (to H.W.) from the National Institutes of Health.

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