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Two crystal forms of 3-carboxy-cis,cis-muconate lactonizing enzyme from Pseudomonas putida have been characterized. Form A is in space group P6, with unit-cell dimensions a = b = 232, c = 79 Å, α = β = 90, γ = 120°. Form B is orthorhombic, with cell dimensions a = 163, b = 139, c = 90 Åα = β = γ = 90°.