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Doc. S1: Carettochelyid peripheral bone histology.

Doc. S2: List of additional trionychid specimens on which the absence of ossicles was tested.

Figure S1. A, B, bone microstructures of peripherals of the carettochelyid turtles Anosteira sp. (FMNH PR 819). C, D, Carettochelys insculpta (MAGNT R12640). E, F, three Pseudanosteira pulchra (UCMP V78031/131731). D, F, images taken in polarized light. A, overview image of the bridge peripheral sampled of Anosteira sp. The dorsal socket for the rib is filled with dark sediment. B, close-up view of the external cortex and interior cancellous bone at the distal tip of the specimen. The compact bone tissue is composed of interwoven structural fibres, whereas the cavities of the cancellous bone are lined with secondary lamellar bone. C, D, close-up view of the external cortical bone and interior cancellous bone. The compact bone, which consists of interwoven structural fibres, is vascularized by reticular primary canals. The bone trabeculae increase in length and are more slender towards the centre of the cancellous bone. E, F, close-up view of the transition from the cancellous bone to the internal compact cortex. The cortex consists of parallel-fibered bone tissue. A small irregularly shaped erosion bay is seen within the outer (more visceral) layers of the compact bone.

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