The HLA dictionary 2008: a summary of HLA-A, -B, -C, -DRB1/3/4/5, and -DQB1 alleles and their association with serologically defined HLA-A, -B, -C, -DR, and -DQ antigens

Abstract

The 2008 report of the human leukocyte antigen (HLA) data dictionary presents serologic equivalents of HLA-A, -B, -C, -DRB1, -DRB3, -DRB4, -DRB5, and -DQB1 alleles. The dictionary is an update of the one published in 2004. The data summarize equivalents obtained by the World Health Organization Nomenclature Committee for Factors of the HLA System, the International Cell Exchange, UCLA, the National Marrow Donor Program, recent publications, and individual laboratories. The 2008 edition includes information on 832 new alleles (685 class I and 147 class II) and updated information on 766 previously listed alleles (577 class I and 189 class II). The tables list the alleles with remarks on the serologic patterns and the equivalents. The serological equivalents are listed as expert assigned types, and the data are useful for identifying potential stem cell donors who were typed by either serology or DNA-based methods. The tables with HLA equivalents are available as a searchable form on the IMGT/HLA database Web site (http://www.ebi.ac.uk/imgt/hla/dictionary.html).

The 2008 edition of the human leukocyte antigen (HLA) data dictionary is the fifth update; previous versions have been published in 1997, 1999, 2001, and 2004 (1–4). The HLA data dictionary is aimed at identifying the serological types associated with each HLA-A, -B, -C, -DRB1, and -DQB1 allelic product. The dictionary is a reference to help with the identification of potential hematopoietic stem cell (HSC) donors. The serologic equivalents of the DNA-defined alleles can be used to identify potential donors listed in unrelated donor registry databases and umbilical cord banks who have been typed by serology. Although the majority of laboratories use DNA-based methods for typing today, there is a significant number of HLA assignments in registries and banks that have been determined by serology-based methods. For example, in the database of almost 7 million volunteers of the National Marrow Donor Program (NMDP) in the United States, the percent of serologic assignments for volunteer donors is still significant, with 45% of the HLA-A and -B loci, 31% of the -C locus, 4% of the -DRB1 locus, and 37% of the -DQB1 locus assignments defined serologically. The availability of serologic equivalents allows potentially matched volunteers typed by serology to be matched to DNA-typed patients.

The number of newly identified HLA alleles grows each year and since the 2004 publication, 832 new HLA alleles have been added, including 235 HLA-A, 310 -B, 140 -C, 116 -DRB1, 9 -DRB3/4/5, and 22 -DQB1 alleles. Information has also been updated for 766 previously listed alleles numbering 158 HLA-A, 303 -B, 116 -C, 133 -DRB1, 13 -DRB3/4/5, and 43 -DQB1 alleles. This edition of the dictionary includes all newly defined four-digit alleles displayed in the IMGT/HLA database (5) to the end of July 2007 in Tables 1–6.

Table 1.  HLA-A alleles and their serologic designations^g

HLA allele	Expert assigned	WHO assigned	International Cell Exchange, UCLA		NMDP		IHWS13 data		NN	Comments
			Assigned type [%]	Cells tested	Assigned type [%]	Cells tested	Assignment	Cells tested	Assigned type >0.55
NT, not tested; NN, neural network; HLA, human leukocyte antigen; WHO, World Health Organization; NMDP, National Marrow Donor Program. a Updated International Cell Exchange, UCLA. b WHO assignment based on dictionary data. c Updated NMDP data. In some cases, the information reported in previous dictionaries has changed. Cells have been excluded from this study in some cases because their HLA typing has changed. d The allele has been identified but serology not informative. e Not assigned; see comment column for predictions >0.20. f If a primary specificity is provided by the NN, other specificities >0.4 are shown in the comments column. ^g  Ref 13–28, 61.
A*0101	A1	A1	A1 [98–100]	34^a	A1 [99]	5612	A1	113	The allele was used for training of the NN – A1
A*01010102N	Not expressed	Null	–	–	–	–	–	–	NT
A*0102	A1	A1	–	–	A1 [95]	129	–	–	Training – A1
A*0103	A1	A1^b	A1 [99]	2	A1 [94]^c	33	A1	5	Training – A1
A*0104N	Not expressed	Null	–	–	^c^d	1	–	–	NT
A*0106	A1	–	–	–	A1 [100]^c	3	–	–	A1
A*0107	A1	A1	–	–	–	–	–	–	A1
A*0108	A1	A1	–	–	–	–	–	–	A1	Normal A1
A*0109	A1	–	–	–	–	–	–	–	A1
A*0110	A1	–	–	–	–	–	–	–	A1
A*0111N	Not expressed	Null	–	–	–	–	–	–	NT
A*0112	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A1
A*0113	A1	–	–	–	–	–	–	–	A1
A*0114	A1	–	–	–	–	–	–	–	A1
A*0115N	Not expressed	Null	–	–	–	–	–	–	NT
A*0116N	Not expressed	Null	–	–	–	–	–	–	NT
A*0117	A1	–	–	–	–	–	–	–	A1
A*0118N	Not expressed	Null	–	–	–	–	–	–	NT
A*0119	A1	A1	–	–	–	–	–	–	Not assigned^e	NN: A1
A*0120	A1	–	–	–	–	–	–	–	A1
A*0121	A1	–	–	–	–	–	–	–	A1
A*0122N	Not expressed	Null	–	–	–	–	–	–	NT
A*0123	A1	–	–	–	–	–	–	–	A1
A*0124	A1	–	–	–	–	–	–	–	A1
A*0125	A1	–	–	–	–	–	–	–	A1
A*0126	A1	–	–	–	–	–	–	–	A1
A*0127N	Not expressed	Null	–	–	–	–	–	–	NT
A*0201	A2	A2	A2 [99–100]	117^a	A2 [98]	6315	A2	230	Training – A2
A*02010102L	Low A2	Low A2	–	–	–	–	–	–	Identical to A*0201	Significantly reduced or low cell surface expression
A*0202	A2	A2	A2 [100]^a	7	A2 [81]	859	A2	8	Training – A2	ihw13: variant serum pattern
A*0203	A203/A2	A203	A2 [99–100]^a	10	A2 [64]	472	A203	12	A2	ihw13: very short A2
A*0204	A2	A2	–	–	A2 [77]	28	–	–	A2
A*0205	A2	A2	A2 [100]	5	A2 [81]	462	A2	10	Training – A2
A*0206	A2	A2	A2 [99–100]^a	47	A2 [68]	636	A2	24	Training – A2
A*0207	A2	A2	A2 [100]^a	15	A2 [80]	135	A2	17	Training – A2
A*0208	A2	A2	–	–	A2 [100]^c	7	–	–	A2
A*0209	A2	A2	–	–	A2 [100]^c	10	–	–	A2
A*0210	A210/A2	A210	–	–	A2 [100]^c	10	–	–	A2
A*0211	A2	A2	A2 [99]	1	A2 [74]	228	A2	17	A2	ihw13: short A2
A*0212	A2	A2	–	–	A2 [100]^c	10	A2	1	A2
A*0213	A2	A2	–	–	A2 [100]^c	17	–	–	A2
A*0214	A2	A2	–	–	A2 [100]^c	7	–	–	A2
A*0215N	Not expressed	Null	–	–	^c	–	–	–	NT
A*0216	A2	A2	–	–	A2 [81]^c	16	–	–	A2
A*0217	A2	A2	–	–	A2 [73]	51	A2	2	A2
A*0218	A2	A2	–	–	A2 [100]^c	1	–	–	A2
A*0219	A2	–	–	–	Blank [100]^c	1	–	–	A2
A*0220	A2	A2	–	–	A2 [100]^c	26	–	–	A2	Normal A2
A*0221	A2	A2	–	–	–	–	–	–	A2
A*0222	A2	A2	–	–	A2 [100]^c	21	–	–	A2
A*0224	A2	A2	–	–	A2 [100]^c	11	–	–	A2
A*0225	A2	A2	–	–	–	–	–	–	A2
A*0226	A2	–	–	–	A2 [100]^c	3	–	–	A2
A*0227	A2	–	–	–	A2 [100]^c	1	–	–	A2
A*0228	A2	–	–	–	A2 [100]^c	1	–	–	A2
A*0229	A2	A2	–	–	A2 [100]^c	2	–	–	A2
A*0230	A2	–	–	–	A2 [100]^c	7	–	–	A2
A*0231	A2	A2	–	–	–	–	–	–	A2
A*0232N	Not expressed	Null	–	–	–	–	–	–	NT
A*0233	A2	–	–	–	A2 [100]^c	3	–	–	A2
A*0234	A2	A2	–	–	–	–	–	–	A2
A*0235	A2	A2	–	–	A2 [100]^c	8	A2	2	A2	Normal A2
A*0236	A2	–	–	–	A2 [100]^c	1	–	–	A2	Normal A2
A*0237	A2	–	–	–	–	–	–	–	A2
A*0238	A2	–	–	–	A2 [67], blank [33]^c	3	–	–	A2	Normal A2
A*0239	A2	–	–	–	A2 [100]^c	1	–	–	A2
A*0240	A2	–	–	–	–	–	–	–	A2
A*0241	A2	A2	–	–	–	–	–	–	A2
A*0242	A2	A2	–	–	–	–	–	–	A2
A*0243N	Not expressed	Null	–	–	–	–	–	–	NT
A*0244	A2/blank	–	–	–	Blank [100]^c	6	–	–	A2
A*0245	A2	–	–	–	A2 [100]^c	2	–	–	A2
A*0246	A2	A2	–	–	–	–	–	–	A2
A*0247	A2	–	–	–	–	–	–	–	A2
A*0248	A2	–	–	–	–	–	–	–	A2	A2x9, few A23 + 24 sera reactive
A*0249	A2	A2	–	–	–	–	A2	1	A2	Normal A2. The HLA club cell exchange: nine laboratories performing serology
A*0250	A2	A2	–	–	–	–	–	–	A2
A*0251	A2	–	–	–	–	–	–	–	A2
A*0252	A2	–	–	–	–	–	–	–	A2
A*0253N	Not expressed	Null	–	–	–	–	–	–	NT
A*0254	A2	–	–	–	–	–	–	–	A2
A*0255	A2/A28	–	–	–	–	–	–	–	Not assigned^e	NN: A2, A28, A69
A*0256	A2	–	–	–	–	–	–	–	A2	Short A2
A*0257	A2	–	–	–	–	–	–	–	A2
A*0258	A2	–	–	–	–	–	–	–	A2
A*0259	A2	A2	–	–	–	–	–	–	A2
A*0260	A2	–	–	–	A2 [100]^c	3	–	–	A2
A*0261	A2	–	–	–	–	–	–	–	A2
A*0262	A2	–	–	–	–	–	–	–	A2	A2 variant; A2x69 variant
A*0263	A2	–	–	–	A2 [100]^c	1	–	–	A2
A*0264	A2	–	–	–	^c^d	1	–	–	A2
A*0265	A2/A19	A2/19	–	–	–	–	–	–	Not assigned^e	NN: A74
A*0266	A2	A2	–	–	A2 [100]^c	2	–	–	A2
A*0267	A2	A2	–	–	–	–	–	–	A2
A*0268	A2	A2	–	–	–	–	–	–	A2
A*0269	A2	–	–	–	–	–	–	–	A2
A*0270	A2	A2	–	–	–	–	–	–	A2
A*0271	A2	A2	–	–	–	–	–	–	A2
A*0272	A2	A2	–	–	–	–	–	–	A2
A*0273	A2	A2	–	–	–	–	–	–	A2
A*0274	A2	A2	–	–	–	–	–	–	A2
A*0275	A2	–	–	–	–	–	–	–	A2
A*0276	A2	–	–	–	–	–	–	–	A2
A*0277	A2	A2	–	–	–	–	–	–	A2
A*0278	Undefined	A2	–	–	–	–	–	–	Not assigned^e	NN: A69, A28
A*0279	A2	A2	–	–	–	–	–	–	A2
A*0280	A2	–	–	–	–	–	–	–	A2
A*0281	A2	–	–	–	A2 [100]^c	1	–	–	A2
A*0282N	Not expressed	Null	–	–	–	–	–	–	NT
A*0283N	Not expressed	Null	–	–	–	–	–	–	NT
A*0284	A2	–	–	–	–	–	–	–	A2
A*0285	A2	–	–	–	A2 [100]^c	1	–	–	A2
A*0286	A2	–	–	–	–	–	–	–	A2
A*0287	A2	–	–	–	–	–	–	–	A2
A*0288N	Not expressed	Null	–	–	–	–	–	–	NT
A*0289	A2	A2	–	–	–	–	–	–	A2
A*0290	A2	A2	–	–	–	–	–	–	A2
A*0291	A2	–	–	–	–	–	–	–	A2
A*0292	A2	–	–	–	–	–	–	–	A2
A*0293	A2	–	–	–	–	–	–	–	A2
A*0294N	Not expressed	Null	–	–	–	–	–	–	NT
A*0295	A2	A2	–	–	–	–	–	–	A2
A*0296	A2	–	–	–	–	–	–	–	A2
A*0297	A2	A2	–	–	–	–	–	–	A2
A*0299	A2	–	–	–	–	–	–	–	A2
A*9201	A2	–	–	–	–	–	–	–	A2
A*9202	A2	–	–	–	–	–	–	–	A2
A*9203	A2	–	–	–	–	–	–	–	A2
A*9204	A2	–	–	–	–	–	–	–	A2
A*9205	A2	A2	–	–	–	–	–	–	A2
A*9206	A2	–	–	–	–	–	–	–	A2
A*9207	A2	–	–	–	–	–	–	–	A2
A*9208	A2	–	–	–	–	–	–	–	A2
A*9209	A2	–	–	–	–	–	–	–	A2
A*9210	A2	–	–	–	–	–	–	–	A2
A*9211	A2	–	–	–	–	–	–	–	A2
A*9212	A2	–	–	–	–	–	–	–	A2
A*9213N	Not expressed	Null	–	–	–	–	–	–	NT
A*9214	A2	–	–	–	–	–	–	–	A2
A*9215	A2	–	–	–	–	–	–	–	A2
A*9216	A2	–	–	–	–	–	–	–	A2
A*9217	A2	–	–	–	–	–	–	–	A2
A*9218	A2	–	–	–	–	–	–	–	A2
A*9219	A2	–	–	–	–	–	–	–	A2
A*9220	A2	–	–	–	–	–	–	–	A2
A*9221	A2	–	–	–	–	–	–	–	A2
A*9222	A2	–	–	–	–	–	–	–	A2
A*9223	A2	–	–	–	–	–	–	–	A2
A*9224	A2	–	–	–	–	–	–	–	A2
A*9225N	Not expressed	Null	–	–	–	–	–	–	NT
A*9226	A2	–	–	–	–	–	–	–	A2
A*0301	A3	A3	A3 [98–100]^a	44	A3 [99]	3504	A3	106	Training – A3
A*03010102N	Not expressed	Null			–	–	–	–	NT
A*0302	A3	A3	A3 [99–100]	2	A3 [78]	342	A3	3	Training – A3
A*0303N	Not expressed	Null	–	–	–	–	–	–	NT
A*0304	A3	A3	–	–	^c^d	1	–	–	A3
A*0305	A3/blank	A3	–	–	Blank [100]^c	5	Blank	1	A3	ihw13: only reactive with some A3 + 11 and A3 + 19 + 34 sera; normal A3
A*0306	A3/blank	–	–	–	–	–	A3/blank	1	A3	ihw13: very short A3
A*0307	A3	–	–	–	A3 [75]^c	4	–	–	A3
A*0308	A3	–	–	–	A3 [100]^c	4	–	–	A3
A*0309	A3	–	–	–	^c^d	1	A3	1	A3
A*0310	A3/blank	–	–	–	–	–	–	–	A3	A3 sera negative, only A3 + 30 + 31 + 33 monoclonal antibodie positive
A*0311N	Not expressed	Null	–	–	–	–	–	–	NT
A*0312	A3	–	–	–	–	–	–	–	A3
A*0313	A3	–	–	–	–	–	–	–	A3
A*0314	A3	–	–	–	–	–	–	–	A3
A*0315	A3	–	–	–	–	–	–	–	A3
A*0316	A3	–	–	–	–	–	–	–	A3
A*0317	A3	–	–	–	–	–	–	–	A3
A*0318	Undefined	–	–	–	–	–	–	–	A1
A*0319	A3	–	–	–	–	–	–	–	A3
A*0320	A3	A3	–	–	–	–	–	–	A3
A*0321N	Not expressed	Null	–	–	–	–	–	–	NT
A*0322	A3	–	–	–	–	–	–	–	A3
A*0323	A3	–	–	–	–	–	–	–	A3
A*0324	A3	A3	–	–	–	–	–	–	A3
A*0325	A3	–	–	–	–	–	–	–	A3
A*0326	A3	–	–	–	–	–	–	–	A3
A*0327	A3	–	–	–	–	–	–	–	A3
A*0328	A3	–	–	–	–	–	–	–	A3
A*0329	A3	–	–	–	–	–	–	–	A3
A*0330	A3	–	–	–	–	–	–	–	A3
A*1101	A11	A11	A11 [99–100]^a	77	A11 [99]	2530	A11	118	Training – A11
A*1102	A11	A11	A11 [99]	1	A11 [76]	42	A11	8	A11	ihw13: short A11
A*1103	A11	A11	–	–	A11 [82]^c	17	A11	1	A11
A*1104	A11	A11	A11 [63–69]	2	A11 [75], blank [25]^c	8	–	–	A11	Short A11; normal A11
A*1105	A11	A11	–	–	A11 [80]^c	5	–	–	A11	Short A11; 3/31 A11 sera reactive
A*1106	A11	–	–	–	A11 [100]^c	2	–	–	A11
A*1107	A11	A11	–	–	–	–	–	–	A11	Normal A11
A*1108	A11	A11	–	–	–	–	–	–	A11	Short A11
A*1109	A11	A11	–	–	A11 [100]^c	1	–	–	A11
A*1110	A11	A11	–	–	Blank [100]^c	1	–	–	A10
A*1111	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A11
A*1112	A11	A11	–	–	A11 [100]^c	3	–	–	A11	Normal A11
A*1113	A11	A11	–	–	–	–	–	–	A11	Short A11
A*1114	A11	A11	–	–	–	–	–	–	A11	Short A11, like A11.2
A*1115	A11	A11	–	–	–	–	–	–	A11
A*1116	A11	–	–	–	–	–	–	–	A11
A*1117	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A11
A*1118	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A11
A*1119	A11	A11	–	–	–	–	–	–	A11
A*1120	A11	–	–	–	–	–	–	–	A11
A*1121N	Not expressed	Null	–	–	–	–	–	–	NT
A*1122	A11	–	–	–	–	–	–	–	A11
A*1123	A11	–	–	–	–	–	–	–	A11
A*1124	A11	–	–	–	–	–	–	–	A11
A*1125	A11	A11	–	–	–	–	–	–	A11
A*1126	A11	–	–	–	–	–	–	–	A11
A*1127	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A11, A1
A*1128	A11	–	–	–	A11 [100]^c	1	–	–	A11
A*1129	A11	–	–	–	–	–	–	–	A11
A*1130	A11	–	–	–	–	–	–	–	A11
A*1131	A11	–	–	–	–	–	–	–	A11
A*1132	A11	–	–	–	–	–	–	–	A11
A*2301	A23	A23(9)	A23 [95–100]^a	29	A23 [95], A24 [1]	5119	A23	42	Training – A9, A23
A*2302	A23	–	–	–	A23 [100]^c	5	–	–	A23
A*2303	A23	–	–	–	A9 [100]^c	1	–	–	A23
A*2304	A23	A23(9)^b	–	–	–	–	A23	1	A23	ihw13: short A23, negative with A9 + 80 (+1) sera
A*2305	A23	–	–	–	A23 [100]^c	2	–	–	A23
A*2306	A23	–	–	–	–	–	–	–	A23
A*2307N	Not expressed	Null	–	–	–	–	–	–	NT
A*2308N	Not expressed	Null	–	–	–	–	–	–	NT
A*2309	A23	–	–	–	–	–	–	–	A23
A*2310	A23	–	–	–	–	–	–	–	A23
A*2311N	Not expressed	Null	–	–	–	–	–	–	NT
A*2312	A23	A23(9)	–	–	–	–	–	–	A23
A*2313	A23	–	–	–	–	–	–	–	A23
A*2314	A23	–	–	–	–	–	–	–	A23
A*2315	A23	–	–	–	–	–	–	–	A23
A*2316	A23	–	–	–	–	–	–	–	A23
A*2402	A24	A24(9)	A24 [97–100]	99^a	A24 [97], A2403 [1]	3098	A24	146	Training – A9, A24
A*24020102L	Low A24	Low A24(9)	A24 [0–5]^a	3	–	–	Blank	1	Identical to A*2402	Significantly reduced or low cell surface expression
A*2403	A2403/A24	A2403	A24 [66–68], A2403 [29–31]^a	10	A24 [55], blank [20], A9 [4], A2403 [1]	282	A24	5	A24	ihw13: short A24, negative with A9 + 80 (+1) sera
A*2404	A24	A24(9)	–	–	A24 [62], blank [25], A2403 [13]^c	8	–	–	A9
A*2405	A9	A9	A23 [67–70], A24 [14–19]^a	2	A24 [45], A23 [50]^c	20	–	–	A24
A*2406	A24	A24(9)	–	–	A24 [67], blank [33]^c	3	A24	5	A24
A*2407	A24	A24(9)	A24 [99–100]^a	21	A24 [59]	320	A24	17	Training – A9, A24
A*2408	A24	A24(9)	–	–	A24 [47], blank [26]	30	–	–	A24	Short A24, similar to A2403
A*2409N	Not expressed	Null	–	–	Blank [100]^c	2	–	–	NT
A*2410	A2403/A24	A2403	–	–	A24 [42], A9 [9], blank [16], A2403 [3]	32	A24	1	A24	A2403; ihw13: short A24
A*2411N	Not expressed	Null	Blank^a	1	Blank [100]^c	2	–	–	NT
A*2413	A24	A24(9)	–	–	^c^d	1	–	–	A24
A*2414	A24	A24(9)	–	–	A24 [82]^c	11	–	–	A24
A*2415	A24	–	–	–	^c	–	–	–	A24
A*2417	A24	A24(9)^b	–	–	A24 [82]^c	11	A24	3	A24	ihw13: short A24
A*2418	A24/A3	A24(9)/A3	–	–	–	–	–	–	A9^f	A2403x3; short A24 with most A3 and A9 reactive; NN: A24
A*2419	A9	A9	–	–	–	–	–	–	A24	Very short A9
A*2420	A24	A24(9)^b	–	–	A24 [100]^c	9	A24	1	A24
A*2421	A24/A9	A24(9)	–	–	–	–	–	–	A24	A9variant; A24 sera negative
A*2422	A24	A9	–	–	A24 [83]^c	6	A24	1	A24	ihw13: short A24, negative with A9 + 80 (+1) sera; A2403 like
A*2423	A24	A2403	–	–	A24 [100]^c	11	–	–	A24	A2403
A*2424	A9	–	–	–	–	–	–	–	A9^f	NN: A23
A*2425	A24	–	–	–	A24 [100]^c	13	–	–	A24
A*2426	A24	–	–	–	A24 [100]^b	2	–	–	A24
A*2427	A24	A24(9)^b	–	–	–	–	A24	1	A24	ihw13: short A24
A*2428	A9	A24(9)	–	–	A24 [50], A9 [50]^c	2	–	–	Not assigned^e	NN: A9, A24, A2
A*2429	A24	–	–	–	A24 [100]^c	1	–	–	A24
A*2430	A24	–	–	–	–	–	–	–	A24
A*2431	A24	–	–	–	–	–	–	–	A24
A*2432	A24	–	–	–	A24 [100]^c	4	–	–	A24
A*2433	A2403/A24	A2403	A24 [99–100]^a	4	A24 [100]^c	4	A24	1	A24	A2403
A*2434	A24	–	–	–	–	–	–	–	A24
A*2435	A24	–	–	–	–	–	–	–	A24
A*2436N	Not expressed	Null	–	–	–	–	–	–	NT
A*2437	A24	A24	A24 [94–97]^a	4	A24 [100]^c	1	–	–	A24
A*2438	A24	–	–	–	–	–	–	–	A24
A*2439	A24	–	–	–	–	–	–	–	A24
A*2440N	Not expressed	Null	–	–	–	–	–	–	NT
A*2441	A24	–	–	–	–	–	–	–	A24
A*2442	A24	–	–	–	–	–	–	–	A24
A*2443	A24	A24(9)	–	–	–	–	–	–	A24
A*2444	A9	–	–	–	–	–	–	–	A9^f	NN: A24
A*2445N	Not expressed	Null	–	–	–	–	–	–	NT
A*2446	A24	–	–	–	–	–	–	–	A24
A*2447	A24	–	–	–	–	–	–	–	A24
A*2448N	Not expressed	Null	–	–	–	–	–	–	NT
A*2449	A24	–	–	–	–	–	–	–	A24
A*2450	A24	–	–	–	–	–	–	–	A24
A*2451	A24	–	–	–	–	–	–	–	A24
A*2452	A24	–	–	–	–	–	–	–	A24
A*2453	A24	–	–	–	–	–	–	–	A24
A*2454	A24	–	–	–	–	–	–	–	A24
A*2455	A24	–	–	–	–	–	–	–	A24
A*2456	A24	–	–	–	–	–	–	–	A24
A*2457	A24	–	–	–	–	–	–	–	A24
A*2458	A24	A24(9)	–	–	–	–	–	–	A24
A*2459	A24	–	–	–	–	–	–	–	A24
A*2460N	Not expressed	Null	–	–	–	–	–	–	NT
A*2461	A24	–	–	–	–	–	–	–	A24
A*2462	A24	–	–	–	–	–	–	–	A24
A*2463	A24	–	–	–	–	–	–	–	A24
A*2464	A24	–	–	–	–	–	–	–	A24
A*2465	A24	–	–	–	–	–	–	–	A24
A*2466	A24	–	–	–	–	–	–	–	A24
A*2467	A24	–	–	–	–	–	–	–	A24
A*2468	A24	–	–	–	–	–	–	–	A24
A*2469	A24	–	–	–	–	–	–	–	A24
A*2470	A24	–	–	–	–	–	–	–	A24
A*2471	A24	–	–	–	–	–	–	–	A24
A*2472	A24	–	–	–	–	–	–	–	A24
A*2473	A24	–	–	–	–	–	–	–	A24
A*2474	A24	–	–	–	–	–	–	–	A24
A*2475	A24	–	–	–	–	–	–	–	A24
A*2476	A24	A24(9)	–	–	–	–	–	–	A24
A*2477	A24	–	–	–	–	–	–	–	A24
A*2478	A24	A24(9)	–	–	–	–	–	–	A24
A*2479	A24	–	–	–	–	–	–	–	A24
A*2501	A25	A25(10)	A25 [94–99]	6^a	A25 [95], A10 [2]	1375	A25	18	Training – A10, A25
A*2502	A25	A25(10)^b	–	–	A25 [88]^c	8	A25	1	A25	Normal A25
A*2503	A25	–	–	–	A25 [100]^c	2	–	–	A25
A*2504	A25	–	–	–	–	–	–	–	A25
A*2505	A25	–	–	–	A2 [100]^c	1			A25
A*2506	A25	–	–	–	–	–			A25
A*2601	A26	A26(10)	A26 [95–97]^a	27	A26 [97], A10 [2]	1730	A26	47	Training – A10, A26
A*2602	A26	A26(10)	A26 [91–97]^a	3	A26 [81]	31	A26	2	Training – A10, A26
A*2603	A26	A26(10)	A26 [92–98]^a	11	A26 [68], A10 [8]	38	A26	5	Training – A10, A26	ihw13: short A26
A*2604	A26	A26(10)	–	–	A10 [50], blank [50]^c	2	–	–	A26
A*2605	A26	A26(10)	–	–	A26 [90], A25 [10]^c	10	–	–	A26
A*2606	A26	A26(10)	–	–	A26 [100]^c	1	–	–	A26
A*2607	A26	A26(10)	–	–	A26 [100]^c	1	–	–	Not asssigned^e	NN: A43, A10, A26
A*2608	A26	A26(10)	–	–	A26 [95]	65	A26	3	A26	ihw13: short A26, similar to A*2603; normal A26
A*2609	A26	A26(10)^b	–	–	A26 [33], A10 [33], A34 [33]^c	3	A26	2	A26	ihw13: short A26
A*2610	A10	A10	–	–	–	–	–	–	A26	Short A26, A10 variant
A*2611N	Not expressed	Null	–	–	–	–	–	–	NT
A*2612	A26	–	–	–	A26 [75], A34 [25]^c	4	–	–	A26
A*2613	A26	–	–	–	A26 [100]^c	3	–	–	A26	NN: A66
A*2614	A26	–	–	–	–	–	–	–	A26
A*2615	A10	A10^b	–	–	–	–	A10	2	A26	ihw13: only few A10 sera reactive, may be typed as blank; A blank
A*2616	A26	–	–	–	A26 [100]^c	1	–	–	A26
A*2617	A26	–	–	–	–	–	–	–	A26
A*2618	A10	–	–	–	Blank [100]^c	1	–	–	A10^f	NN: A26
A*2619	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A43, A10
A*2620	A26	–	–	–	–	–	–	–	A26
A*2621	A26	–	–	–	–	–	–	–	A26
A*2622	A26	A26(10)	–	–	–	–	–	–	A26
A*2623	A26	A26(10)	–	–	–	–	–	–	A26
A*2624	A26	–	–	–	–	–	–	–	A26
A*2625N	Not expressed	Null	–	–	–	–	–	–	NT
A*2626	A26	A10	–	–	–	–	–	–	A26
A*2627	A26	–	–	–	–	–	–	–	A26
A*2628	A26	–	–	–	–	–	–	–	A26
A*2629	A26	A26(10)	–	–	–	–	–	–	A26
A*2630	A26	–	–	–	–	–	–	–	A26
A*2631	A26	–	–	–	–	–	–	–	A26
A*2632	A26	–	–	–	–	–	–	–	A26
A*2633	A26	–	–	–	–	–	–	–	A26
A*2634	A26	–	–	–	–	–	–	–	A26
A*2635	A26	–	–	–	–	–	–	–	A26
A*2901	A29	A29(19)	A29 [98]^a	1	A29 [96]	115	A29	10	Training – A29
A*29010102N	Not expressed	Null	–	–	–	–	–	–	NT
A*2902	A29	A29(19)	A29 [98–100]^a	17	A29 [99]	1488	A29	28	Training – A29
A*2903	A29	–	–	–	A29 [100]^c	4	–	–	A29
A*2904	A29	–	–	–	A29 [100]^c	1	–	–	A29
A*2905	A29	–	–	–	–	–	–	–	A29
A*2906	A29	–	–	–	–	–	–	–	A29
A*2907	A29	–	–	–	–	–	–	–	A29
A*2908N	Not expressed	Null	–	–	–	–	–	–	NT
A*2909	A29	–	–	–	–	–	–	–	A29
A*2910	A29	A29(19)	–	–	A29 [100]^c	2	–	–	A29
A*2911	A29	–	–	–	–	–	–	–	A29
A*2912	A29	–	–	–	–	–	–	–	A29
A*2913	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A32, A29
A*2914	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A29, A30
A*2915	A29	–	–	–	–	–	–	–	A29
A*2916	A29	A29(19)	–	–	–	–	–	–	A29
A*3001	A30	A30(19)	A30 [97–100]^a	25	A30 [89], A31 [3], blank [3], A19 [1]	2895	A30	28	Training – A30
A*3002	A30	A30(19)	A30 [96–100]^a	17	A30 [93], A31 [2], A19 [1]	2124	A30	25	Training – A30
A*3003	A30	A30(19)	–	–	A30 [83], blank [17]^c	6	–	–	A30
A*3004	A30	A30(19)	A30 [97–98]^a	2	A30 [94]	85	A30	7	Training – A30
A*3006	A30	–	–	–	^c^d	1	–	–	A30
A*3007	A30	–	–	–	A30 [100]^c	2	–	–	A30
A*3008	A30	–	–	–	–	–	–	–	A30
A*3009	A30	–	–	–	A30 [100]^c	3	A30	1	A30	ihw13: very short A30, only broad A19 sera reactive
A*3010	A30	–	–	–	A30 [100]^c	10	–	–	A30
A*3011	A30	A30(19)	–	–	–	–	–	–	A30
A*3012	A30	–	–	–	–	–	–	–	A30
A*3013	A30	–	–	–	–	–	–	–	A30
A*3014L	A30	Low A30(19)	–	–	–	–	–	–	A30	Significantly reduced or low cell surface expression
A*3015	A30	A30(19)	–	–	–	–	–	–	A30
A*3016	A30	–	–	–	–	–	–	–	A30
A*3017	A30	A30(19)	–	–	–	–	–	–	A30
A*3018	A30	A30(19)	–	–	–	–	–	–	A30
A*3019	A30	–	–	–	–	–	–	–	A30
A*3020	A30	–	–	–	–	–	–	–	A30
A*3021	A30	–	–	–	–	–	–	–	A30
A*3022	A30	–	–	–	–	–	–	–	A30
A*3101	A31	A31(19)	A31 [94–98]^a	23	A31 [90], A30 [3], A19 [1]	3517	A31	58	Training – A31
A*3102	A31	A31(19)^b	–	–	A31 [100]^c	17	–	–	A31
A*3103	A31	–	–	–	A31 [100]^c	2	–	–	A31
A*3104	A31	A31(19)	–	–	–	–	–	–	A31
A*3105	A31	A31(19)	–	–	–	–	–	–	A31
A*3106	A31	–	–	–	–	–	–	–	A31
A*3107	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A32, A23, A9
A*3108	A31/A24	–	–	–	A32 [100]^c	1	–	–	Not assigned^e	A31x24; previously A*2416; NN: A23, A9, A32
A*3109	A31	–	–	–	A31 [100]^c	3	–	–	A31	Bw4 motif present
A*3110	A31	A31(19)	–	–	–	–	–	–	Not assigned^e	NN: A32
A*3111	A31	–	–	–	–	–	–	–	A31
A*3112	A31	A31(19)	–	–	–	–	–	–	A31
A*3113	A31	–	–	–	–	–	–	–	A31
A*3114N	Not expressed	Null	–	–	–	–	–	–	NT
A*3115	A31	–	–	–	–	–	–	–	A31
A*3116	A31	–	–	–	–	–	–	–	A31
A*3117	A31	–	–	–	–	–	–	–	A31
A*3118	A31	–	–	–	–	–	–	–	A31
A*3201	A32	A32(19)	A32 [97–99]^a	20	A32 [94]	3441	A32	50	Training – A32
A*3202	A32	A32(19)	–	–	A32 [100]^c	3	–	–	Not assigned^e	NN: A32
A*3203	A32	–	–	–	A32 [67], blank [33]^c	3	–	–	A32
A*3204	A32/A3	–	–	–	A3 [75], A19 [13], A32 [13]^c	8	A‘3204’	1	Not assigned^e	A3 with most A32 sera reactive (ihw13); NN: A32, A3
A*3205	A32	–	–	–	–	–	–	–	A32
A*3206	A32	–	–	–	–	–	–	–	A32
A*3207	A32	–	–	–	A32 [100]^c	1	–	–	A32
A*3208	A32	–	–	–	A32 [100]^c	2	–	–	A32
A*3209	A32	–	–	–	–	–	–	–	A32
A*3210	A32	–	–	–	–	–	–	–	A32
A*3211Q	A32	–	–	–	–	–	–	–	A32	Not expected to be expressed
A*3212	A32	–	–	–	–	–	–	–	A32
A*3213	A32	–	–	–	A32 [100]^c	1	–	–	A32
A*3214	A32	A32(19)	–	–	–	–	–	–	A32
A*3215	A32	–	–	–	–	–	–	–	A32
A*3301	A33	A33(19)	A33 [97–100]	10^a	A33 [87], A19 [3], blank [4]	687	A33	19	Training – A33
A*3303	A33	A33(19)	A33 [95–100]^a	36	A33 [95], A34 [1]	887	A33	27	Training – A33
A*3304	A33	–	–	–	A33 [50], blank [50]^c	2	–	–	A33
A*3305	A33	A33(19)	–	–	A33 [92]^c	13	A33	1	A33	ihw13: short A33
A*3306	A33	–	–	–	^c^d	1	–	–	A33
A*3307	A33	–	–	–	–	–	–	–	A33
A*3308	Undefined	A33(19)	–	–	–	–	–	–	Not assigned^e	NN: A31, A33
A*3309	Undefined	A19	–	–	–	–	–	–	Not assigned^e	NN: A31, A33
A*3310	A33	–	–	–	–	–	–	–	A33
A*3311	A33	–	–	–	–	–	–	–	A33
A*3312	A33	–	–	–	–	–	–	–	A33
A*3313	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: A33
A*3401	A34	A34(10)	A34 [93–98]^a	30	A34 [67], A26 [10], A10 [5], A66 [4], A33 [2], blank [7]	269	A34	25	Training – A10, A34	ihw13: distinct pattern from A*3402
A*3402	A34	A34(10)	A34 [92–95]^a	11	A34 [89], A33 [4], A26 [3], A10 [2]	996	A34	11	Training – A10, A34	ihw13: distinct pattern from A*3401
A*3403	A34	–	–	–	–	–	–	–	A34
A*3404	A34	A34(10)	–	–	–	–	A34	2	A34	ihw13: short A34, with some A11 and A19 sera reactive; A34 variant with some A10 sera reactive
A*3405	A34	–	–	–	–	–	–	–	A34
A*3406	A10	–	–	–	–	–			A10^f	NN: A34
A*3407	A34	–	–	–	–	–			A34
A*3408	A34	–	–	–	A10 [100]^c	1			A34
A*3601	A36	A36	A36 [85–93], A1 [5–8]^a	5	A36 [68], A1 [16], blank [6]	807	A36	7	Training – A36
A*3602	A36	–	–	–	–	–	–	–	A36
A*3603	A36	A36	–	–	–	–	–	–	A36
A*3604	A36	A36^b	–	–	–	–	A36	2	A36
A*4301	A43	A43	–	–	A26 [50], A10 [13], A34 [13], A19 [13], blank [13]^c	8	–	–	Training – A43
A*6601	A66	A66(10)	A66 [37–59], A‘6601’ [22–42], A26 [15–18], A10 [2–7]^a	14	A26 [61], A66 [20], A34 [8], A10 [3], A25 [1]	799	A66	19	Training – A10, A66	ihw13: A‘6601’ pattern – different from A‘6602’; reactive with A34 + 66 sera
A*6602	A66	A66(10)	A‘6602’ [35–65], A66 [15–42], A34 [7–18], A10 [4]^a	8	A66 [38], A34 [27], A10 [10], A74 [5], A26 [4]	205	A66	2	A66	ihw13: A‘6602’, short A66
A*6603	A10	A10	–	–	A66 [32], A34 [18], A10 [7], A26 [7]	28	–	–	A10^f	Short A10; NN: A26
A*6604	A66	–	–	–	–	–	–	–	A66
A*6605	A26	–	–	–	A26 [100]^c	1	–	–	A26
A*6606	A66	A66(10)	–	–	–	–	–	–	A66
A*6801	A68	A68(28)	A68 [53–68], A28 [31–45]^a	34	A28 [63], A68 [27]	2736	A68	49	Training – A28, A68
A*6802	A68	A68(28)	A68 [55–65], A28 [32–43]^a	15	A28 [72], A68 [19], A69 [1]	2327	A68	18	Training – A28, A68
A*6803	A68	A68(28)^b	A68 [47–66], A28 [34–48]^a	16	A28 [63], A68 [28]	202	A68	4	A68
A*6804	A68	A68(28)	–	–	A28 [75], A68 [25]^c	4	–	–	A68
A*6805	A68	A68(28)	–	–	A28 [53], A68 [29]	59	A28	1	A68	ihw13: possibly A68, but cell also A2 positive
A*6806	A68	–	–	–	A28 [50], A68 [50]^c	2	–	–	A68
A*6807	A68	–	–	–	Blank [100]^c	1	–	–	A68
A*6808	A68	A68(28)	A68 [38–45], A28 [53]	2	A28 [100]^c	2	–	–	A68
A*6809	A68	–	–	–	A68 [100]^c	1	–	–	A68
A*6810	A68	A28^b	–	–	Blank [50], A1 [50]^c	2	A28	1	A68	ihw13: possibly A68, but cell also A2 positive
A*6811N	Not expressed	Null	–	–	–	–	–	–	NT
A*6812	A28	A28	–	–	A28 [50], blank [50]^c	6	–	–	A68	Short A28
A*6813	A68	–	–	–	–	–	–	–	A68
A*6814	A68	–	–	–	–	–	–	–	A68
A*6815	A68	–	–	–	–	–	–	–	A68
A*6816	A68	A68(28)	–	–	–	–	–	–	A68
A*6817	A68	A28^b	–	–	A28 [100]^c	1	A28	1	A68	ihw13: possibly A68, but cell also A2 positvie
A*6818N	Not expressed	Null	–	–	–	–	–	–	NT
A*6819	A68	–	–	–	A28 [100]^c	1	–	–	A68
A*6820	A68	–	–	–	–	–	–	–	A68
A*6821	A68	–	–	–	–	–	–	–	A68
A*6822	A68	–	–	–	–	–	–	–	A68
A*6823	A68	–	–	–	A68 [100]^c	1	–	–	A68
A*6824	A68	–	–	–	–	–	–	–	A68
A*6825	A68	–	–	–	A28 [100]^c	1	–	–	A68
A*6826	A68	A28	–	–	–	–	–	–	A68
A*6827	A68	–	–	–	–	–	–	–	A68
A*6828	A68	–	–	–	–	–	–	–	A68
A*6829	A68	–	–	–	–	–	–	–	A68
A*6830	A68	–	–	–	–	–	–	–	A68
A*6831	A68	–	–	–	–	–	–	–	A68
A*6832	A68	–	–	–	–	–	–	–	A68
A*6833	A68	A68(28)	–	–	–	–	–	–	A68
A*6834	A68	–	–	–	–	–	–	–	A68
A*6835	A68	–	–	–	–	–	–	–	A68
A*6836	A68	A68(28)	–	–	–	–	–	–	A28^f	NN: A68
A*6837	A68	–	–	–	–	–	–	–	A68
A*6838	A68	–	–	–	–	–	–	–	A68
A*6839	A68	–	–	–	–	–	–	–	A68
A*6840	A68	–	–	–	–	–	–	–	A68
A*6901	A69	A69(28)	A69 [40–70], A28 [33–49]^a	3	A28 [56], A69 [7], A2 [6], A68 [3]	289	A69	6	Training – A28, A69
A*7401	A74	A74(19)	A74 [72–95]^a	13	A74 [40], A19 [11], blank [36]	115	A74	11	Training – A74
A*7402	A74	A74(19)	–	–	Blank [50], A74 [17], A33 [17], A31 [17]^c	6	–	–	A74
A*7403	A74	A74(19)	A74 [89–91]	5	Blank [72], A74 [17]	29	A74	4	A74
A*7404	A74	–	–	–	A74 [100]^c	1	–	–	A74
A*7405	A74	–	–	–	Blank [100]^c	1	–	–	A74
A*7406	A74	A74(19)	–	–	A74 [100]^c	1	A74	1	A74
A*7407	A74	A74(19)	–	–	–	–	–	–	A74
A*7408	A74	–	–	–	–	–	–	–	A74
A*7409	A74	–	–	–	A74 [33], A33 [33], blank [33]^c	3	–	–	A74
A*7410	Undefined	–	–	–	–	–			Not assigned^e	NN: A74
A*7411	A74	–	–	–	^c^d	1			A74
A*7412N	Not expressed	Null	–	–	–	–			NT
A*8001	A80	A80	A80 [68–88], A36 [0–3]	11	Blank [77], A80 [10], A36 [2], A1 [1]	505	A80	9	Training – A80	ihw13: difficult to type next to A1 or A24

Table 2.  HLA-B alleles and their serologic designations^g

HLA allele	Expert assigned	WHO assigned	International Cell Exchange, UCLA		NMDP		IHWS13 data		NN	Comments
			Assigned type [%]	Cells tested	Assigned type [%]	Cells tested	Assignment	Cells tested	Assigned type >0.55
NT, not tested; NN, neural network; HLA, human leukocyte antigen; WHO, World Health Organization; NMDP, National Marrow Donor Program. a Updated International Cell Exchange, UCLA. b Updated NMDP data. In some cases the information reported in previous dictionaries has changed. Cells have been excluded from this study in some cases because their HLA typing has changed. c WHO assignment based on dictionary data. d The allele has been identified but serology not informative. e Not assigned; see comment column for predictions >0.20. f If a primary specificity is provided by the NN, other specificities >0.4 are shown in the comments column. ^g  Ref 13–16, 29–57, 62.
B*0702	B7	B7	B7 [98–100]^a	32	B7 [98]	10,841	B7	83	Training – B7
B*0703	B703/B7	B703	–	–	B7 [94]^b	17	B703, B7, blank	1	B7	Short B7, B7 + 42 sera reactive, B7 + 22 sera negative
B*0704	B7	B7	–	–	B7 [89]	44	B7	2	Training – B7
B*0705	B7	B7	B7 [91–97]^a	4	B7 [95]	42	B7	6	Training – B7
B*0706	B7	B7	B7 [94–95]	4	B7 [96]	23	–	–	B7
B*0707	B7	B7	–	–	B7 [93]^b	15	–	–	B7
B*0708	B7	B7^c	–	–	–	–	B7	1	B7	Short B7, negative with B7 + 22 sera, similar to B703
B*0709	B7	B7	B7 [97–98]	2	B7 [100]^b	8	B7	2	B7
B*0710	B7	B7^c	–	–	B7 [92]^b	12	B7	2	B7
B*0711	B7	B7	–	–	–	–	–	–	B7
B*0712	B7	B7^c	–	–	B7 [100]^b	5	–	–	B7
B*0713	B7/blank	–	–	–	Blank [100]^b	3	–	–	B7
B*0714	B7	B7^c	–	–	B7 [100]^b	10	B7	1	B7
B*0715	B7	B7	–	–	B7 [100]^b	1	–	–	B7	B7 variant, some Bw6 sera negative
B*0716	B7	B7	–	–	B7 [50], B60 [50]^b	2	–	–	B7	Short B7, few B7 sera reactive, similar to B703
B*0717	B7	B7	–	–	–	–	–	–	B7
B*0718	B7	–	–	–	–	–	–	–	B7
B*0719	B7/42	–	–	–	B42 [100]^b	1	–	–	B42	Short B7
B*0720	B7/blank	–	–	–	–	–	–	–	B7	Short B7, B7 sera negative, B7 + 42 sera reactive
B*0721	B7	–	–	–	B7 [100]^b	1	–	–	B7
B*0722	B7	–	–	–	–	–	–	–	B7
B*0723	B7	–	–	–	^b^d	1	–	–	B7
B*0724	B7	B7	–	–	–	–	–	–	B7	B7 weakly expressed, reactive with B7 + 27 (+40) sera
B*0725	B7	–	–	–	–	–	–	–	B7
B*0726	B7	B7	–	–	B7 [100]^b	1	–	–	B7	Normal B7
B*0727	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B7
B*0728	B7	–	–	–	–	–	–	–	B7
B*0729	B7	–	–	–	–	–	–	–	B7
B*0730	B7	B7	–	–	–	–	–	–	B7	B7 variant
B*0731	B7/B42	B7/B42	–	–	–	–	–	–	B7	B7x42, short B7
B*0732	B7/blank	–	–	–	–	–	–	–	B7	B blank
B*0733	B7	–	–	–	B7 [100]^b	1	–	–	B7
B*0734	B7	–	–	–	–	–	–	–	B7
B*0735	B7	–	–	–	–	–	–	–	B7
B*0736	B7	–	–	–	B7 [100]^b	1	–	–	B7	Bw4 positive
B*0737	B7	–	–	–	–	–	–	–	B7
B*0738	Undefined	–	–	–	–	–	–	–	Not assigned^e
B*0739	B7	–	–	–	–	–	–	–	B7
B*0740	B7	B7	–	–	–	–	–	–	B7
B*0741	B7	–	–	–	–	–	–	–	B7
B*0742	B7	–	–	–	–	–	–	–	B7
B*0743	B7	–	–	–	–	–	–	–	B7
B*0744	B7	–	–	–	–	–	–	–	B7
B*0745	B7	–	–	–	–	–	–	–	B7
B*0746	B7	B7	–	–	–	–	–	–	B7
B*0747	B7	–	–	–	–	–	–	–	B7
B*0748	B7	–	–	–	–	–	–	–	B7
B*0749N	Not expressed	Null	–	–	–	–	–	–	NT
B*0750	Undefined	–	–	–	–	–	–	–	NT	NN: B7
B*0751	B7	–	–	–	–	–	–	–	B7
B*0752	B7	–	–	–	–	–	–	–	B7
B*0753	B7	–	–	–	–	–	–	–	B7
B*0754	B7	–	–	–	–	–	–	–	B7
B*0755	B7	–	–	–	–	–	–	–	B7
B*0756	B7	–	–	–	–	–	–	–	B7
B*0757	B7	B7	–	–	–	–	–	–	B7
B*0758	B7	B7	–	–	–	–	–	–	B7
B*0801	B8	B8	B8 [100]^a	9	B8 [99]	6518	B8	61	Training – B8
B*0802	B8	B8	–	–	^b^d	2	B‘0802’	1	Not assigned^e	B8 Bw4; ihw13: B8 Bw4 with extra reactions; NN: B8
B*0803	B8	B8	–	–	B8 [100]^b	1	–	–	Not assigned^e	B8 Bw4
B*0804	B8/blank	–	B8 [51–71], B59 [6–7]	3	B8 [80], blank [20]^b	5	B8	1	B8	Blank Bw6; ihw13: short B8
B*0805	B8	–	–	–	^b^d	1	–	–	B8
B*0806	B8	B8	–	–	B8 [100]^b	1	–	–	B8
B*0807	B8	B8	–	–	B8 [100]^b	1	–	–	B8
B*0808N	Not expressed	Null	–	–	–	–	–	–	NT
B*0809	B8	B8	–	–	B8 [100]^b	11	B8	2	B8
B*0810	B8	B8	–	–	–	–	–	–	B8	Short B8
B*0811	B8	–	–	–	B8 [100]^b	3	–	–	B8
B*0812	B8	–	–	–	^b^d	1	–	–	B8
B*0813	B8	–	–	–	B8 [100]^b	1	–	–	B8
B*0814	B8	–	–	–	–	–	–	–	B8
B*0815	B8	B8	–	–	–	–	B8	1	B8
B*0816	B8	B8	–	–	–	–	–	–	B8
B*0817	Undefined	–	–	–	–	–	–	–	Not assigned^e	B blank; NN: B8
B*0818	B8	B8	–	–	–	–	–	–	B8
B*0819N	Not expressed	Null	–	–	–	–	–	–	NT
B*0820	B8	–	–	–	–	–	–	–	B8
B*0821	B8	–	–	–	–	–	–	–	B8
B*0822	B8	–	–	–	–	–	–	–	B8
B*0823	B8	–	–	–	–	–	–	–	B8
B*0824	B8	–	–	–	–	–	–	–	B8
B*0825	B8	–	–	–	–	–	–	–	B8
B*0826	B8	–	–	–	–	–	–	–	B8
B*0827	B8	–	–	–	–	–	–	–	B8
B*0828	B8	–	–	–	–	–	–	–	B8
B*0829	B8	B8	–	–	–	–	–	–	B8
B*0830N	Not expressed	Null	–	–	–	–	–	–	NT
B*0831	B8	–	–	–	–	–	–	–	B8
B*0832	B8	–	–	–	–	–	–	–	B8
B*0833	B8	–	–	–	–	–	–	–	B8
B*1301	B13	B13	B13 [95–99]^a	4	B13 [97]	238	B13	9	Training – B13
B*1302	B13	B13	B13 [98–100]^a	9	B13 [99]	1307	B13	17	Training – B13
B*1303	B49/B15	–	–	–	B13 [100]^b	1	–	–	B13	B15x21–Bw4; B21 variant with some B15 sera reactive
B*1304	B49/B15	B15/B21	B21 [22–38], B49 [15], B15 [7–17], B15x21 [0–15], B50 [5–11], B63 [0–8], B77 [4–7]	2	–	–	B‘1304’	1	B13	B15x21–Bw4, similar to B*1303; ihw13: new pattern, also B49, B63 and B77 reactivity
B*1306	B13	–	–	–	–	–	–	–	B13
B*1307N	Not expressed	Null	–	–	–	–	–	–	NT
B*1308	B13	–	–	–	–	–	–	–	B13
B*1309	Undefined	–	–	–	–	–	–	–	Not assigned^e	Bw6pos; NN: B13
B*1310	B13	–	–	–	B13 [100]^b	2	–	–	Not assigned^e	NN: B13 B15
B*1311	B13	–	–	–	B13 [100]^b	1	–	–	B13
B*1312	B13	–	–	–	–	–	–	–	B13
B*1313	B13	B13			–	–	–	–	B13
B*1314	B13	–	–	–	–	–	–	–	B13
B*1315	B13	–	–	–	–	–	–	–	B13
B*1316	B13	–	–	–	–	–	–	–	B13
B*1317	B13	–	–	–	–	–	–	–	B13
B*1318	B13	–	–	–	–	–	–	–	B13
B*1319	B13	–	–	–	–	–	–	–	B13
B*1320	B13	–	–	–	–	–	–	–	B13
B*1401	B64	B64(14)	B14 [49–58], B64 [32–43], B65 [2–10]^a	8	B14 [72], B64 [10], B65 [10]	1003	B64	11	Training – B14
B*1402	B65	B65(14)	B65 [50–54], B14 [45–47], B64 [1–3]^a	9	B14 [71], B65 [25], B64 [2]	2244	B65	22	Training – B14
B*1403	B14	B14^c	–	–	B14 [50], B65 [35]	40	–	–	B14
B*1404	B14	–	–	–	B14 [67], B64 [33]^b	3	–	–	B14
B*1405	B14	–	–	–	B14 [40], blank [60]^b	5	B65	1	B14
B*1406	B14	B14	–	–	B14 [100]^b	8	–	–	B14	B65
B*1407N	Not expressed	Null	–	–	–	–			NT
B*1501	B62	B62(15)	B62 [93–98]^a	30	B62 [96], B15 [2]	1906	B62	64	Training – B15 B62
B*15010102N	Not expressed	Null	–	–	–	–	–	–	NT
B*1502	B75	B75(15)	B75 [79–86], B15 [8–11], B62 [5–12]	16	B75 [62], B62 [22], B15 [7]	1035	B75	18	Training – B15 B75
B*1503	B72	B72(70)	B70 [47–61], B72 [31–45]^a	17	B70 [61], B72 [10], B35 [2], B62 [1], blank [11]	2569	B72	25	Training – B70 B72
B*1504	B62	B62(15)	B62 [90], B15 [5], B75 [4]^a	1	B62 [69], B15 [10]	29	B62	1	B62
B*1505	B62	B62(15)	B62 [88]	1	B62 [64], B15 [18]	22	B62	3	B62
B*1506	B62	B62(15)	–	–	B62 [81]^b	13	–	–	B62
B*1507	B62	B62(15)	B62 [93]	2	B62 [79]	101	B62	2	Training – B15 B62
B*1508	B75/B62	B75(15)	B75 [30–48], B15 [19–35], B62 [6–19], B‘1508’ [11–14]	2	B62 [20], B15 [18], B75 [18]	45	B75	3	B15
B*1509	B70	B70	–	–	B70 [53], blank [33]	65	B71	2	Training – B70 B71
B*1510	B71	B71(70)	B70 [41–49], B71 [41–47]^a	15	B70 [49], B35 [4], B72 [2], B71 [2], B75 [1], B15 [1], blank [13]	1204	B71	7	Training – B70 B71
B*1511	B75	B75(15)	B75 [31–51], B15 [21–38], B46 [6–12], B‘1511’ [0–19]	3	B62 [27], B75 [21], B46 [15], B15 [15]	68	B75	4	B15
B*1512	B76	B76(15)	B76 [76–88], B45 [3–6]	5	B76 [29], B62 [18], B75 [6], B15 [29]^b	17	B76	2	Training – B15 B76
B*1513	B77	B77(15)	B77 [46–53], B63 [0–16]	2	B77 [21], B63 [19], B15 [15], B75 [12], blank [13], B62 [8], B53 [6]	116	B77	7	Training – B15 B77
B*1514	B76	B76(15)	B76 [23], B45 [20], B15 [20], B70 [6], B63 [5]	1	B45 [80], B44 [20]^b	5	B76	1	B62
B*1515	B75/62	B62(15)	B75 [43–57], B62 [31–42], B15 [8–15]^a	10	B62 [49], B75 [28], B15 [13]	138	B75	1	B15	ihw13: short B75
B*1516	B63	B63(15)	B63 [96–97]^a	8	B63 [80], B57 [3], B15 [3]	816	B63	3	Training – B15 B63
B*1517	B63	B63(15)	B63 [92–96]	5	B63 [88], B15 [3], B57 [1]	797	B63	11	Training – B15 B63
B*1518	B71	B71(70)	B70 [52–64], B71 [28–34]^a	11	B70 [51], blank [34], B71 [3]	552	B71	8	Training – B70 B71
B*1519	B76	B76(15)	B76 [29], B15 [19], B45 [11], B62 [8], B75 [7]	1	B70 [50], B76 [50]^b	2	–	–	B76
B*1520	B62	B62(15)	B62 [73], B15 [18], B75 [5]	1	B62 [100]^b	10	B62	1	B62	ihw13: short B62
B*1521	B75	B75(15)	B75 [80–88], B15 [6–9], B62 [3–10]	21	B75 [55], B62 [12], B15 [7]	132	B75	9	Training – B15 B75
B*1523	B70/B5/blank	–	B‘1523’ [17], B5 [14–15], B77 [7–11], B70 [7–9], B15 [5–8]	2	Blank [50], B70 [17], B51 [17], B53 [8]^b	12	B‘1523’	2	Not assigned^e	B5x70–Bw4, B5/53/77 variant; NM5; ihw13: new pattern, short B71-Bw4; NN: B70 B71
B*1524	B62	B62(15)	B62 [45–48], B‘1524’ [23–24], B15 [21–22], B77 [4–6], B63 [2–3]	4	B62 [44], blank [22]	54	B62	8	B15	B62 Bw4
B*1525	B62	B62(15)	B62 [86–94], B75 [2–10]^a	10	B62 [74], B75 [8], B15 [7]	116	B62	1	Training – B15 B62	ihw13: short B62, short B75
B*1526N	Not expressed	Null	–	–	–	–	–	–	NT
B*1527	B62	B62(15)	B62 [97]^a	1	B62 [76], B15 [7]	76	B62	2	Training – B15 B62	ihw13: short B62, short B75
B*1528	B62	B62(15)^c	–	–	B62 [80], B15 [20]^b	5	–	–	B62
B*1529	B70	B15	B70 [54–58], B15 [11–12], B71 [7–10]	2	Blank [71], B70 [19]	21	B71	2	B71
B*1530	B62	B62(15)^c	B62 [63–86], B75 [4–13], B15 [12–19]	3	B62 [64], B15 [17]	98	B62	3	Training – B15 B62	ihw13: short B62, short B75
B*1531	B75	B75(15)	–	–	B75 [38], B62 [38]^b	13	B75	1	B75	ihw13: short B75
B*1532	B62	B62(15)	–	–	B62 [100]^b	5	–	–	B62
B*1533	B62	B15	–	–	–	–	–	–	B62	B62
B*1534	B62	B62(15)^c	–	–	B62 [100]^b	8	–	–	B62	Normal B62
B*1535	B62	B62(15)^c	B62 [86–93], B75 [5–7]^a	14	B62 [79]	34	B62	5	Training – B15 B62
B*1536	Undefined	–	–	–	B13 [100]^b	3	–	–	B15	Bw4 positive, NN: B77
B*1537	B70	B70^c	B70 [46], B71 [11], B78 [9]	2	Blank [54], B70 [30]	24	–	–	B71
B*1538	B62	–	–	–	B62 [100]^b	3	–	–	B62	B62 variant, also reactive with B52 sera
B*1539	B62	B62(15)^c	B62 [92–98]	2	B62 [84], B15 [13]^b	38	B62	3	B62
B*1540	B62/blank	–	–	–	Blank [61], B70 [10], B62 [8]	26	–	–	B62	Short B62
B*1542	B15	–	–	–	–	–	–	–	B62	Short B15; reactive with some B22 sera
B*1543	B15	–	–	–	–	–	–	–	B15^f	Bw4 positive, NN: B62
B*1544	B75	–	–	–	–	–	–	–	B75
B*1545	B62	B62(15)	–	–	B62 [92]^b	13	–	–	B62
B*1546	Undefined	B72(70)	–	–	B50 [50], B72 [25]^b	4	–	–	Not assigned^e	NN: B15 B62
B*1547	B70/blank	–	–	–	Blank [57], B35 [14]^b	14	–	–	B72	B48, all B15 sera negative
B*1548	B62	B62(15)	–	–	B62 [100]^b	3	–	–	B62
B*1549	B72	–	–	–	–	–	–	–	B72
B*1550	B62	–	–	–	–	–	–	–	B62
B*1551	B70	B70	–	–	B35 [100]^b	1	–	–	B71
B*1552	Undefined	B15	–	–	Blank [67]^b	3	–	–	B71	B15
B*1553	Undefined	–	–	–	–	–	–	–	Not assigned^e
B*1554	B72	–	–	–	B70 [50], B72 [50]^b	2	–	–	B72
B*1555	B15	B15	–	–	–	–	–	–	B15^f	B15 Bw6; NN: B75
B*1556	B62	–	–	–	–	–	–	–	B62
B*1557	B62	–	–	–	–	–	–	–	B62
B*1558	B62	B62(15)	–	–	B62 [100]^b	3	–	–	B62	B15 or B62
B*1560	B62	–	–	–	–	–	–	–	B62
B*1561	B72	–	–	–	–	–	–	–	B72
B*1562	Undefined	–	–	–	B62 [67]^b	3	–	–	Not assigned^e	NN: B70 B72
B*1563	B62	–	–	–	B62 [100]^b	2	–	–	B62
B*1564	B71	–	–	–	–	–	–	–	B71
B*1565	B62	–	–	–	B62 [100]^b	2	–	–	B62
B*1566	B62	–	–	–	B62 [100]^b	1	–	–	B62
B*1567	B63	–	–	–	–	–	–	–	B63	Bw4 positive
B*1568	Undefined	B35	–	–	–	–	–	–	Not assigned^e	NN: B70 B72
B*1569	B72	–	–	–	–	–	–	–	B72
B*1570	B62	B62(15)	–	–	–	–	B62	1	B62	ihw13: short B62, short B75
B*1571	B62	B62(15)	–	–	B62 [83], B75 [17]^b	6	–	–	B62
B*1572	B71	–	–	–	–	–	–	–	B71
B*1573	B62	B62(15)	–	–	B62 [100]^b	1	–	–	B62
B*1574	B72	–	–	–	–	–	–	–	B72
B*1575	B62	–	–	–	B62 [100]^b	1	–	–	B62
B*1576	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B15
B*1577	B62	–	–	–	–	–	–	–	B62
B*1578	B62	B15	–	–	–	–	–	–	B62
B*1579N	Not expressed	Null	–	–	–	–	–	–	NT
B*1580	B70	B70	–	–	–	–	–	–	B71	B70
B*1581	B62	–	–	–	–	–	–	–	B62
B*1582	B62	B62(15)	–	–	–	–	–	–	B62	B62 + A30/31
B*1583	B62	–	–	–	–	–	–	–	B62	NN: B15
B*1584	B62	B62(15)	–	–	–	–	–	–	B62
B*1585	B62	–	–	–	–	–	–	–	B62
B*1586	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B15, B62
B*1587	B15	–	–	–	–	–	–	–	B15
B*1588	B75	–	–	–	–	–	–	–	B75
B*1589	B77	–	–	–	–	–	–	–	B77
B*1590	B71	–	–	–	–	–	–	–	B71
B*1591	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B70, B72, B12
B*1592	B62	–	–	–	–	–	–	–	B62
B*1593	B71	B71(70)	–	–	–	–	–	–	B71
B*1594N	Not expressed	Null	–	–	–	–	–	–	NT
B*1595	B63	–	–	–	–	–	–	–	B63
B*1596	B62	B62(15)	–	–	–	–	–	–	B62
B*1597	B62	–	–	–	–	–	–	–	B62
B*1598	B72	–	–	–	–	–	–	–	B72
B*1599	B71	–	–	–	–	–	–	–	B71
B*9501	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B15, B46
B*9502	B62	B62(15)	–	–	–	–	–	–	B62
B*9503	B70	B70	–	–	–	–	–	–	B72
B*9504	B62	–	–	–	–	–	–	–	B62
B*9505	B62	–	–	–	–	–	–	–	B62
B*9506	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B15, B62
B*9507	B62	B62(15)	–	–	–	–	–	–	B62
B*9508	B71	B71(70)	–	–	–	–	–	–	B71
B*9509	B62	–	–	–	–	–	–	–	B62
B*9510	B62	–	–	–	–	–	–	–	B62
B*9511N	Not expressed	Null	–	–	–	–	–	–	NT
B*9512	B75	B15	–	–	–	–	–	–	B75
B*9513	B62	–	–	–	–	–	–	–	B62
B*9514	B70	B70	–	–	–	–	–	–	B71
B*9515	B70	–	–	–	–	–	–	–	B70
B*9516	B62	–	–	–	–	–	–	–	B62
B*9517	B62	–	–	–	–	–	–	–	B62
B*9518	B62	–	–	–	–	–	–	–	B62
B*9519	B71	–	–	–	–	–	–	–	B71
B*9520	B62	–	–	–	–	–	–	–	B62
B*9521	B75	–	–	–	–	–	–	–	B75
B*9522	B62	–	–	–	–	–	–	–	B62
B*9523	B72	–	–	–	–	–	–	–	B72
B*9524	B71	–	–	–	–	–	–	–	B71
B*9525	B62	–	–	–	–	–	–	–	B62
B*9526	B62	–	–	–	–	–	–	–	B62
B*9527	B72	–	–	–	–	–	–	–	B72
B*9528	B62	–	–	–	–	–	–	–	B62
B*9529	B62	–	–	–	–	–	–	–	B62
B*9530	B62	–	–	–	–	–	–	–	B62
B*9532	B72	–	–	–	–	–	–	–	B72
B*1801	B18	B18	B18 [97–100]^a	12	B18 [96]	1641	B18	53	Training – B18
B*1802	B18	B18	–	–	B18 [90]	31	B18	3	Training – B18
B*1803	B18	B18	–	–	B18 [83], blank [17]	29	B18	1	B18
B*1804	B18	B18^c	–	–	B18 [84], blank [11]^b	19	B18	1	Not assigned^e	NN: B18
B*1805	B18	B18	–	–	B18 [83]^b	6	B18	1	B18	ihw13: short B18, blank
B*1806	B18	B18	–	–	–	–	–	–	B18	B18 variant, negative with Bw4 and Bw6 sera
B*1807	B18/blank	–	–	–	Blank [100]^b	1	B18	1	B18	ihw13: short B18, blank, B18 spec sera negative
B*1808	B18	–	–	–	B18 [100]^b	1	–	–	B18
B*1809	B18	B18	–	–	–	–	–	–	Not assigned^e	B18, Bw4; NN: B18, B38
B*1810	B18	–	–	–	–	–	–	–	B18
B*1811	B18	–	–	–	–	–	–	–	B18
B*1812	B18	–	–	–	B18 [50], blank [50]^b	2	–	–	B18
B*1813	B18	–	–	–	B18 [100]^b	2	–	–	B18
B*1814	B18	–	–	–	B18 [100]^b	2	–	–	B18
B*1815	B18	–	–	–	B18 [100]^b	1	–	–	B18
B*1817N	Not expressed	Null	–	–	–	–	–	–	NT
B*1818	B18	–	–	–	B18 [100]^b	4	–	–	B18
B*1819	B18	–	–	–	–	–	–	–	B18
B*1820	B18	–	–	–	–	–	–	–	B18
B*1821	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B18
B*1822	B18	–	–	–	–	–	–	–	B18
B*1823N	Not expressed	Null	–	–	–	–	–	–	NT
B*1824	B18	–	–	–	–	–	–	–	B18
B*1825	B18	–	–	–	–	–	–	–	B18
B*1826	B18	–	–	–	–	–	–	–	B18
B*2701	B27	B27	–	–	B27 [100]^b	12	–	–	B27
B*2702	B27	B27	B27 [97]	1	B27 [87]	417	B27	9	Training – B27	ihw13: short B27
B*2703	B27	B27	B27 [97]	1	B27 [88]	26	B27	5	Training – B27
B*2704	B27	B27	B27 [98]^a	1	B27 [97]	31	B27	7	B27
B*2705	B27	B27	B27 [99–100]	6	B27 [99]	1194	B27	22	Training – B27
B*2706	B27	B27	B27 [97–99]^a	10	B27 [97]	29	B27	6	B27	Previously also named B*2722
B*2707	B27	B27	–	–	B27 [100]^b	49	B27	5	B27
B*2708	B2708/B27/B7	B2708	B2708 [36–58], B7 [19–41], B27 [11–19]	3	B27 [31], B7 [34], blank [16], B2708 [9]^b	32	B2708	1	B27	B7 variant, most B27 sera negative; ihw13: also B73 B7 blank
B*2709	B27	B27	B27 [100]	1	B27 [100]^b	15	–	–	B27
B*2710	B27	B27	–	–	B27 [100]^b	5	–	–	B27
B*2711	B27	B27	–	–	B27 [100]^b	2	–	–	Not assigned^e	B27 variant, extra reactivity with B40 sera; NN: B27
B*2712	Undefined	B27	–	–	Blank [78], B60 [9], B27 [4], B61 [4]^b	23	Blank	1	Not assigned^e	Blank Bw6; B27 sera negative, B7 + 27 (+40) sera positive; ihw13: blank, B61 B27–Bw6; NN: B27
B*2713	B27	B27	–	–	–	–	–	–	B27
B*2714	B27	B27^c	–	–	B27 [100]^b	14	–	–	B27
B*2715	B27	–	–	–	–	–	–	–	B27	Blank Bw4
B*2716	B27	–	–	–	–	–	–	–	B27	Short B27, some B40 sera reactive
B*2717	B27	B27	–	–	–	–	–	–	B27	Normal B27
B*2718	Undefined	–	–	–	Blank [100]^b	2	–	–	Not assigned^e	Bw6 positive
B*2719	B27	B27	–	–	–	–	–	–	B27	Normal B27
B*2720	B27	B27	–	–	–	–	–	–	B27
B*2721	B27	–	–	–	–	–	–	–	B27
B*2723	Undefined	–	–	–	–	–	–	–	Not assigned^e	Blank Bw6, Bw4 negative/weak, only reactive with few B44 + 27 sera
B*2724	B27	–	–	–	–	–	–	–	B27
B*2725	B27	–	–	–	–	–	–	–	B27
B*2726	B27	–	–	–	–	–	–	–	B27
B*2727	B27	–	–	–	–	–	–	–	B27
B*2728	B27	–	–	–	–	–	–	–	B27
B*2729	B27	B27	–	–	–	–	–	–	Not assigned^e	NN: B27
B*2730	B27	B27	–	–	–	–	–	–	B27
B*2731	B27	–	–	–	–	–	–	–	B27
B*2732	B27	–	–	–	–	–	–	–	B27
B*2733	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B27, B40, B61
B*2734	B27	–	–	–	–	–	–	–	B27	NN: B27
B*2735	B27	–	–	–	–	–	–	–	B27
B*2736	B27	–	–	–	–	–	–	–	B27
B*2737	B27	–	–	–	–	–	–	–	B27
B*2738	B27	B27	–	–	–	–	–	–	B27
B*3501	B35	B35	B35 [97–100]^a	27	B35 [98]	2023	B35	58	Training – B35
B*3502	B35	B35	B35 [94–99]	3	B35 [72]	1013	B35	9	Training – B35
B*3503	B35	B35	B35 [95–100]	3	B35 [94]	1226	B35	22	Training – B35
B*3504	B35	B35	–	–	B35 [100]	26	–	–	B35
B*3505	B35	B35	B35 [98–99]	9	B35 [94]	231	B35	7	Training – B35
B*3506	B35	B35	–	–	B35 [100]^b	13	–	–	B35
B*3507	B35	B35	–	–	–	–	–	–	B35
B*3508	B35	B35	B35 [95–98]^a	6	B35 [91]	345	B35	5	Training – B35
B*3509	B35	B35	–	–	B35 [100]^b	3	–	–	B35
B*3510	B35	B35^c	–	–	B35 [77], B70 [15]^b	13	B35	1	B35	ihw13: short B35
B*3511	B35	B35	–	–	B35 [88]^b	16	–	–	B35
B*3512	B35	B35	B35 [98–100]^a	12	B35 [78]	422	B35	2	B35
B*3513	B35	B35	–	–	–	^b	–	–	B35
B*3514	B35	B35	B35 [98]^a	1	B35 [79]	29	–	–	B35
B*3515	B35	B35	–	–	Blank [50], B35 [33]^b	6	–	–	B35	Short B35, B75 CREG sera reactive
B*3516	B35	B35^c	B35 [63], B70 [20]	2	B35 [33], blank [34]	41	B35	1	B35	ihw13: short B35
B*3517	B35	B35	B35 [97–100]	9	B35 [93]	261	B35	2	Training – B35
B*3518	B35	B35	–	–	B35 [100]^b	1	–	–	B35
B*3519	B35/blank	B35	–	–	B35 [40], B70 [20], B77 [20]^b	5	Blank	2	B35	ihw13: new pattern, blank, very short B35
B*3520	B35	B35	–	–	B35 [42], blank [13]	23	B35	5	B35
B*3521	B35/B78	–	–	–	B35 [42], B78 [33], blank [25]^b	12	B78	1	B35	ihw13: B35/78 variant
B*3522	B35	–	–	–	B35 [100]^b	2	–	–	B35
B*3523	B35	B35^c	B35 [84–88]	4	B35 [92]^b	13	B35	1	B35
B*3524	B35	–	–	–	B35 [75], B78 [25]^b	8	–	–	B35
B*3525	B35	B35^c	–	–	B35 [33], blank [67]^b	3	B35	1	B35	ihw13: short B35
B*3526	B35	–	–	–	–	–	–	–	B35
B*3527	B35	B35	–	–	B35 [100]^b	2	–	–	B35
B*3528	B35	–	–	–	B35 [50], blank [43]^b	14	–	–	B35
B*3529	B35	B35	–	–	B35 [100]^b	2	–	–	B35
B*3530	B35	B35	–	–	B35 [100]^b	7	–	–	B35	Normal B35
B*3531	B35	–	–	–	Blank [67], B61 [17], B48 [17]^b	6	B35	1	B35	ihw13: short B35; reactive with few B40 and B35 sera, Korean
B*3532	B35	B35	–	–	B35 [100]^b	2	–	–	B35
B*3533	B35	–	–	–	–	–	–	–	B35
B*3534	B35	–	–	–	B35 [100]^b	1	–	–	B35
B*3535	B35	B35	–	–	–	–	–	–	B35
B*3536	B35	–	–	–	B35 [100]^b	1	–	–	B35
B*3537	B35	–	–	–	B35 [100]^b	1	–	–	B35
B*3538	B35	–	–	–	^b^d	1	–	–	B35
B*3539	B35	–	–	–	–	–	–	–	B35
B*3540N	Not expressed	Null	–	–	–	–	–	–	NT
B*3541	B35	B35	–	–	B35 [100]^b	3	–	–	B35
B*3542	B35	B35	–	–	–	–	–	–	B35	Normal B35
B*3543	B35	B35	B35 [95–97]^a	3	B35 [63], B70 [12]	379	B35	2	Not assigned^e	Previously B*1522; NN: B35
B*3544	Undefined	B35	–	–	–	–	–	–	Not assigned^e	Previously B*1559
B*3545	B35	–	–	–	–	–	–	–	B35
B*3546	B35	B35	–	–	–	–	–	–	B35
B*3547	B35	–	–	–	B35 [100]^b	1	–	–	B35
B*3548	B35	–	–	–	–	–	–	–	B35
B*3549	B35	–	–	–	B35 [100]^b	1	–	–	B35
B*3550	B35	B35	–	–	–	–	–	–	B35
B*3551	B35	B35	–	–	–	–	–	–	B35
B*3552	B35	B35	–	–	–	–	–	–	B35
B*3553N	Not expressed	Null	–	–	–	–	–	–	NT
B*3554	B35	–	–	–	–	–	–	–	B35
B*3555	B35	B35	–	–	–	–	–	–	B35
B*3556	B35	–	–	–	–	–	–	–	B35
B*3557	B35	B35	–	–	–	–	–	–	B35
B*3558	B35	–	–	–	–	–	–	–	B35
B*3559	B35	–	–	–	–	–	–	–	B35
B*3560	B35	B35	–	–	–	–	–	–	Not assigned^e	NN: B35
B*3561	B35	B35	–	–	–	–	–	–	B35
B*3562	B35	–	–	–	–	–	–	–	B35
B*3563	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B40
B*3564	B35	–	–	–	–	–	–	–	B35
B*3565Q	B35	–	–	–	–	–	–	–	B35	Not expected to be expressed
B*3566	B35	–	–	–	Blank [100]^b	1	–	–	B35
B*3567	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B35
B*3568	B35	–	–	–	–	–	–	–	B35
B*3569	B35	–	–	–	–	–	–	–	B35
B*3570	B35	B35	–	–	–	–	–	–	B35
B*3571	B35	–	–	–	–	–	–	–	B35
B*3572	B35	–	–	–	–	–	–	–	B35
B*3573	B35	–	–	–	–	–	–	–	B35
B*3574	B35	–	–	–	–	–	–	–	B35
B*3575	B35	–	–	–	–	–	–	–	B35
B*3576	B35/22	B35/22	–	–	–	–	–	–	B22^f	NN: B56
B*3577	B35	–	–	–	–	–	–	–	B35
B*3701	B37	B37	B37 [98–100]^a	6	B37 [93]	1721	B37	27	Training – B37
B*3702	B37/B27	–	–	–	B27 [33], blank [33], B37 [17], B47 [17]^b	6	–	–	Not assigned^e	B37x27 or blank; few B37 and few B27 sera reactive; NN: B37
B*3703N	Not expressed	Null	–	–	–	–	–	–	NT
B*3704	B37	B37^c	–	–	B37 [50], blank [50]^b	2	B37	1	B37	B37 variant, reactive with few B37 and B18 sera
B*3705	Blank	–	–	–	–	–	–	–	Not assigned^e	Blank Bw6, B37, and Bw4 sera negative, Korean; NN: B37
B*3706	B37	–	–	–	–	–	–	–	B37
B*3707	Undefined	–	–	–	–	–	–	–	Not assigned^e
B*3708	B37	B37	–	–	–	–	–	–	B37
B*3709	B37	B37	–	–	–	–	–	–	Not assigned^e	NN: B37
B*3710	B37	B37	–	–	–	–	–	–	B37
B*3711	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B37
B*3712	B37	–	–	–	–	–	–	–	B37
B*3713	B37	–	–	–	–	–	–	–	B37
B*3801	B38	B38(16)	B38 [96–100]^a	14	B38 [94], B16 [1], B39 [1]	2317	B38	43	Training – B16 B38
B*3802	B38	B38(16)	B38 [91–99]^a	15	B38 [87], B39 [2], B16 [2]	788	B38	15	Training – B16 B38	ihw13: different from B*3801
B*3803	B38	B16	–	–	Blank [100]^b	1	–	–	Not assigned^e	B16 Bw4, B38, and B39 sera negative; NN: B16 B38
B*3804	B38	–	–	–	–	–	–	–	B38
B*3805	B38	B38(16)	–	–	–	–	–	–	B38	Normal B38
B*3806	B38	–	–	–	–	–	–	–	B38
B*3807	B38	–	–	–	–	–	–	–	B38
B*3808	B38	–	–	–	–	–	–	–	B38
B*3809	B38	B16^c	–	–	–	–	–	–	B38	B16–Bw4, on cell VOA Ter 299
B*3810	B38	–	–	–	–	–	–	–	B38
B*3811	B38	–	–	–	–	–	–	–	B38
B*3812	B38	B38(16)	–	–	–	–	–	–	B38
B*3813	B38	–	–	–	–	–	–	–	B38
B*3814	B38	–	–	–	–	–	–	–	B38
B*3815	B38	B38(16)	–	–	–	–	–	–	B38
B*3816	B38	–	–	–	–	–	–	–	B38
B*3901	B3901/B39	B3901	B39 [96–99]^a	7	B39 [95]	476	B39	21	Training – B16 B39
B*39010102L	Low B39	Low B39			–	–	–	–	Identical to B*3901
B*3902	B3902/B39	B3902	B39 [97–99]^a	7	B39 [87]	96	B39	5	Not assigned^e	ihw13: short B39; NN: B16 B39
B*3903	B39	B39(16)	–	–	B39 [89]	19	B39	1	B39
B*3904	B39	B39(16)	–	–	–	^b	–	–	B39
B*3905	B39	B39^c	B39 [89–92], B16 [4–7], B38 [1–5]^a	17	B39 [82]	245	B39	4	B39	B39 variant, ST-16; ihw13: B39 and B38 sera positive
B*3906	B39	B39(16)	B39 [96–99]	6	B39 [94]	508	B39	8	Training – B16 B39
B*3907	B16	–	–	–	Blank [100]^b	1	–	–	B16
B*3908	B39	B39(16)	B39 [86–91], B16 [3–5]	4	B39 [48], blank [41]	57	B39	1	Not assigned^e	ihw13: B39 and B38 sera positive; NN: B16
B*3909	B39	B39(16)	B39 [96–99]	6	B39 [100]^b	29	B39	2	B39
B*3910	B39	B39(16)	B39 [98]^a	4	B39 [82], blank [16]	127	B39	5	B39
B*3911	B39	B39(16)^c	–	–	B39 [79], blank [14]^b	29	–	–	B39
B*3912	B39	B39(16)	–	–	B39 [100]^b	1	B39	1	B39
B*3913	B39	B39(16)	–	–	B39 [40], blank [40]^b	5	–	–	Not assigned^e	NN: B16, B39
B*3914	B39	–	–	–	B39 [100]^b	4	–	–	B39
B*3915	B39	B39(16)^c	–	–	B39 [100]^b	6	B39	1	B39
B*3916	B39	–	–	–	–	–	–	–	B39
B*3917	B39	–	–	–	B39 [100]^b	1	–	–	Not assigned^e	NN: B39, B16
B*3918	B39	–	–	–	–	–	–	–	B39
B*3919	B39	B39(16)^c	–	–	Blank [100]^b	1	B39	1	B39
B*3920	B39	–	B39 [73], B16 [16]^a	1	Blank [67], B38 [33]^b	3	–	–	B16^f	NN: B38
B*3922	B39	–	–	–	^b^d	1	–	–	B39
B*3923	B39	B39(16)	–	–	–	–	–	–	Not assigned^e	NN: B16, B39
B*3924	B39	B39(16)	–	–	B39 [97]^b	30	B39	1	B39	Normal B39
B*3925N	Not expressed	Null	–	–	–	–	–	–	NT
B*3926	B39	–	–	–	–	–	–	–	B39
B*3927	B39	B39(16)	–	–	–	–	–	–	B39	Normal B39
B*3928	B39	–	–	–	–	–	–	–	B39
B*3929	B39	–	–	–	–	–	–	–	B39
B*3930	B39	–	–	–	–	–	–	–	B39
B*3931	B39	–	–	–	B39 [100]^b	2	–	–	B39
B*3932	B16	–	–	–	–	–	–	–	B16^f	NN: B39
B*3933	B39	–	–	–	–	–	–	–	B39
B*3934	B39	B39(16)	–	–	–	–	–	–	B39
B*3935	B39	–	–	–	B39 [100]^b	1	–	–	B39
B*3936	Undefined	–	–	–	–	–	–	–	Not assigned^e	NN: B39, B16, B14
B*3937	B39	–	–	–	–	–	–	–	B39
B*3938Q	B39	–	–	–	–	–	–	–	B39	Not expected to be expressed
B*3939	B39	–	–	–	–	–	–	–	B39
B*3940N	Not expressed	Null	–	–	–	–	–	–	NT
B*3941	B39	B39(16)	–	–	–	–	–	–	B39
B*3942	B39	–	–	–	–	–	–	–	B39
B*4001	B60	B60(40)	B60 [96–98]^a	37	B60 [93], B40 [4], B61 [2]	3854	B60	48	Training – B40 B60
B*4002	B61	B61(40)	B61 [75–95], B40 [3–19], B60 [2–21]^a	24	B61 [80], B60 [8], B40 [6]	1460	B61	33	Training – B40 B61
B*4003	B61	B61(40)^c	–	–	B61 [62], B60 [23]	31	B61	1	B61
B*4004	B61	B61(40)^c	&ndas