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VIII. REFERENCES

  • Abril, A. & Bucher, E. H. (1999). The effects of overgrazing on soil microbial community and fertility in the Chaco dry savannas of Argentina. Applied Soil Ecology 12, 159167.
  • Akashi, N. & Nakashizuka, T. (1999). Effects of bark-stripping by sika deer (Cervus nippon) on population dynamics of a mixed forest in Japan. Forest Ecology and Management 113, 7582.
  • Allard, V., Soussana, J., Falcimagne, R., Berbigier, P., Bonnefond, J., Ceschia, E., D’hour, P., Hénault, C., Laville, P., Martin, C. & Pinarès-patino, C. (2007). The role of grazing management for the net biome productivity and greenhouse gas budget (CO2, N2O and CH4) of semi-natural grassland. Agriculture, Ecosystems and Environment 121, 4758.
  • Andreae, M. O. & Merlet, P. (2001). Emission of trace gases and aerosols from biomass burning. Global Biogeochemical Cycles 15, 955966.
  • Asner, G. P., Elmore, A. J., Olander, L. P., Martin, R. E. & Harris, A. T. (2004). Grazing systems, ecosystem responses, and global change. Annual Review of Environment and Resources 29, 261299.
  • Asner, G. P., Levick, S. R., Kennedy-Bowdoin, T., Knapp, D. E., Emerson, R., Jacobson, J., Colgan, M. S. & Martin, R. E. (2009). Large-scale impacts of herbivores on the structural diversity of African savannas. Proceedings of the National Academy of Sciences 106, 49474952.
  • Atjay, G. L., Ketner, P. & Duvigneaud, P. (1979). Terrestrial primary production and phytomass. In The global carbon cycle (eds B. Bolin, E. T., Degens, S. Kempe & P. Ketner), pp. 129181. John Wiley & Sons, Chichester.
  • Augustine, D. J. & McNaughton, S. J. (2004). Regulation of shrub dynamics by native browsing ungulates on East African rangeland. Journal of Applied Ecology 41, 4558.
  • Ayres, E., Heath, J., Possell, M., Black, H. I. J., Kerstiens, G. & Bardgett, R.D. (2004). Tree physiological responses to above-ground herbivory directly modify below-ground processes of soil carbon and nitrogen cycling. Ecology Letters 7, 469479.
  • Bakker, E. S., Olff, H., Boekhoff, M., Gleichman, J. M. & Berendse, F. (2004). Impact of herbivores on nitrogen cycling: contrasting effects of small and large species. Oecologia 138, 91101.
  • Bakker, E. S., Olff, H. & Gleichman, J. M. (2009). Contrasting effects of large herbivore grazing on smaller herbivores. Basic and Applied Ecology 10, 141150.
  • Baldi, R., Albon, S. D. & Elston, D. A. (2001). Guanacos and sheep: evidence for continuing competition in arid Patagonia. Oecologia 129, 561570.
  • Bardgett, R. D., Frankland, J. C. & Whittaker, J. B. (1993). The effects of agricultural practices on the soil biota of some upland grasslands. Agriculture, Ecosystems and Environment 45, 2545.
  • Bardgett, R. D. & Wardle, D. A. (2003). Herbivore-mediated linkages between aboveground and belowground communities. Ecology 84, 22582268.
  • Bardgett, R. D., Wardle, D. A. & Yeates, G. W. (1998). Linking above-ground and below-ground interactions: how plant responses to foliar herbivory influence soil organisms. Soil Biology and Biochemistry 30, 18671878.
  • Barford, C. C., Wofsy, S. C., Goulden, M. L., Munger, J. W., Pyle, E. H., Urbanski, S. P., Hutyra, L., Saleska, S. R., Fitzjarrald, D. & Moore, K. (2001). Factors controlling long- and short-term sequestration of atmospheric CO2 in a mid-latitude forest. Science 294, 16881691.
  • Bazely, D. R. & Jefferies, R. L. (1985). Goose feces: a source of nitrogen for plant growth in a grazed salt-marsh. Journal of Applied Ecology 22, 693703.
  • Bee, J. N., Kunstler, G. & Coomes, D. A. (2007). Resistance and resilience of New Zealand tree species to browsing. Journal of Ecology 95, 10141026
  • Bellingham, P. J. & Allan, C. N. (2003). Forest regeneration and the influences of white-tailed deer (Odocoileus virginianus) in cool temperate New Zealand rain forests. Forest Ecology and Management 175, 7186.
  • Belovsky, G. E. (1997). Optimal foraging and community structure: the allometry of herbivore food selection and competition. Evolutionary Ecology 11, 641672.
  • Belsky, A. J. (1986). Does herbivory benefit plants? A review of the evidence. American Naturalist 127, 870892.
  • Belsky, A. J. (1987). The effects of grazing: confounding of ecosystem, community, and organism scales. American Naturalist 129, 777783.
  • Belsky, A. J. & Blumenthal, D. M. (1997). Effects of livestock grazing on stand dynamics and soils in upland forests of the Interior West. Conservation Biology 11, 315327.
  • Belsky, A. J., Carson, W. P., Jensen, C. L. & Fox, G. A. (1993). Overcompensation by plants: herbivore optimization or red herring? Evolutionary Ecology 7, 109121.
  • Berendse, F., Breemen, N. V., Rydin, H., Buttler, A., Heijmans, M., Hoosbeek, M. R., Lee, J. A., Mitchell, E., Saarinen, T., Vasander, H. & Wallén, B. (2001). Raised atmospheric CO2 levels and increased N deposition cause shifts in plant species composition and production in Sphagnum bogs. Global Change Biology 7, 591598.
  • Berg, W. A., Bradford, J. A. & Sims, P. L. (1997). Long-term soil nitrogen and vegetation change on Sandhill Rangeland. Journal of Range Management 50, 482486.
  • Bergman, M. (2002). Can saliva from moose, Alces alces, affect growth responses in the sallow, Salix caprea? Oikos 96,164168.
  • Beschta, R. L. & Ripple, W. J. (2008). Wolves, trophic cascades, and rivers in the Olympic National Park, USA. Ecohydrology 1, 118130.
  • Beschta, R. L. & Ripple, W. J. (2009). Large predators and trophic cascades in terrestrial ecosystems of the western United States. Biological Conservation 142, 24012414.
  • Bodmer, R. & Ward, D. (2006). Frugivory in large mammalian herbivores. In Large herbivore ecology, ecosystem dynamics and conservation. (eds K. Danell, R. Bergström, P. Duncan & J. Pastor), pp 232260. Cambridge University Press, Cambridge.
  • Bodmer, R. E. (1989). Ungulate biomass in relation to feeding strategy within Amazonian forests. Oecologia 81, 547550.
  • Bond, W. J. (2005). Large parts of the world are brown or black: a different view on the ‘Green World’ hypothesis. Journal of Vegetation Science 16, 261266.
  • Bond, W. J. & Keeley, J. E. (2005). Fire as a global ‘herbivore’: the ecology and evolution of flammable ecosystems. Trends in Ecology & Evolution 20, 387394.
  • Brand, M. D. & Goetz, H. (1986). Vegetation of exclosures in southwestern North Dakota. Journal of Range Management 39, 434437.
  • Brashares, J. S., Arcese, P., Sam, M. K., Coppolillo, P. B., Sinclair, A. R. E. & Balmford, A. (2004). Bushmeat hunting, wildlife declines, and fish supply in West Africa. Science 306, 11801183.
  • Brathen, K. D., Ims, R. A., Yoccoz, N. G., Fauchald, P., Tveraa, T. & Hausner, V. H. (2007). Induced shift in ecosystem productivity? Extensive scale effects of abundant large herbivores. Ecosystems 10, 773789.
  • Briggs, J. M., Knapp, A. K., Blair, J. M., Heisler, J. L., Hoch, G. A., Lett, M. S. & McCarron, J. K. (2005). An ecosystem in transition: causes and consequences of the conversion of mesic grassland to shrubland. BioScience 55, 243254.
  • Brown, J. H. & Heske, E. J. (1990). Control of a desert-grassland transition by a keystone rodent guild. Science 250, 17051707.
  • Bryant, J. P. & Reichart, P. B. (1992). Controls over secondary metabolite production in arctic woody plants. In Arctic ecosystems in a changing climate: an ecophysiological perspective. (eds F. S. Chapin, R. L. Jefferies, J. F. Reynolds, G. R. Shaver & J. Svoboda), pp. 377390. Academic Press, New York.
  • Bryant, N. A., Johnson, L. F., Brazel, A. J., Balling, R. C., Hutchinson, C. F. & Beck, L. R. (1990). Measuring the effect of overgrazing in the Sonoran Desert. Climatic Change 17, 243264.
  • Bucher, E. H. (1987). Herbivory in arid and semi-arid regions of Argentina. Revista Chilena de Historia Natural 60, 265273.
  • Burney, D. A. & Flannery, T. F. (2005). Fifty millennia of catastrophic extinctions after human contact. Trends in Ecology & Evolution 20, 395401.
  • Butler, L. G. & Kielland, K. (2008). Acceleration of vegetation turnover and element cycling by mammalian herbivory in riparian ecosystems. Journal of Ecology 96, 136144.
  • Calenge, C., Maillard, D., Gaillard, J., Merlot, L. & Peltier, R. (2002). Elephant damage to trees of wooded savanna in Zakouma National Park, Chad. Journal of Tropical Ecology 18, 599614.
  • Cao, G. M., Tang, Y. H., Mo, W. H., Wang, Y. S., Li, Y. N. & Zhao, X. Q. (2004). Grazing intensity alters soil respiration in an alpine meadow on the Tibetan plateau. Soil Biology and Biochemistry 36, 237243.
  • Carter, A. J. & Scholes, R. J. (2000). Spatial global database of soil properties. International Geosphere-Biosphere Programme Data Information Systems. Toulouse.
  • Caspersen, J. P., Pacala, S. W., Jenkins, J. C., Hurtt, G. C., Moorcroft, P. R. & Birdsey, R. A. (2000). Contributions of land-use history to carbon accumulation in U.S. forests. Science 290, 114851.
  • Caughley, G. (1970). Eruption of ungulate populations, with emphasis on Himalayan Thar in New Zealand. Ecology 51, 5372.
  • Chapman, S. K., Hart, S. C., Cobb, N. S., Whitham, T. G. & Koch, G. W. (2003). Insect herbivory increases litter quality and decomposition: an extension of the acceleration hypothesis. Ecology 84, 28672876.
  • Chaneton, E. J. & Lavado, R. S. (1996). Soil nutrients and salinity after long-term grazing exclusion in a flooding Pampa grassland. Journal of Range Management 49, 182187.
  • Classen, A., Overby, S., Hart, S., Koch, G. & Whitham, T. (2007). Season mediates herbivore effects on litter and soil microbial abundance and activity in a semi-arid woodland. Plant and Soil 295, 217227.
  • Coetsee, C., Bond, W. J. & February, E. C. (2010). Frequent fire affects soil nitrogen and carbon in an African savanna by changing woody cover. Oecologia 162, 10271034.
  • Coley, P. D. & Barone, J. A. (1996). Herbivory and plant defenses in tropical forests. Annual Review of Ecology and Systematics 27, 305335.
  • Coley, P. D., Bryant, J. P. & Chapin, F. S. (1985). Resource availability and plant antiherbivore defense. Science 230, 895899.
  • Committee on Climate Change [CCC]. (2008). Building a low-carbon economy—the UK's contribution to tackling climate change. The Stationery Office, Norwich.
  • Conant, R. T., Paustian, K. & Elliott, E. T. (2001). Grassland management and conversion into grassland: effects on soil carbon. Ecological Applications 11, 343355.
  • Coomes, D. A. & Allen, R. B. (2007). Mortality and tree-size distributions in natural mixed-age forests. Journal of Ecology 95, 2740.
  • Coomes, D. A., Allen, R. B., Forsyth, D. M. & Lee, W. G. (2003). Factors preventing the recovery of New Zealand forests following control of invasive deer. Conservation Biology 17, 450459.
  • Coomes, D. A., Allen, R. B., Scott, N. A., Goulding, C. & Beets, P. (2002). Designing systems to monitor carbon stocks in forests and shrublands. Forest Ecology and Management 164, 89108.
  • Cornelissen, J. H. C., Quested, H. M., Gwynn-Jones, D., Van Logtestijn, R. S. P., De Beus, M. A. H., Kondratchuk, A., Callaghan, T. V. & Aerts, R. (2004). Leaf digestibility and litter decomposability are related in a wide range of subarctic plant species and types. Functional Ecology 18, 779786.
  • Côté, S. D., Rooney, T. P., Tremblay, J. P., Dussault, C. & Waller, D. M. (2004). Ecological impacts of deer overabundance. Annual Review of Ecology, Evolution, and Systematics 35, 113147.
  • Coughenour, M. B. (1991). Biomass and nitrogen responses to grazing of upland steppe on Yellowstone's northern winter range. Journal of Applied Ecology 28, 7182.
  • Coughenour, M. B., McNaughton, S. J. & Wallace, L. L. (1985). Shoot growth and morphometric analyses of Serengeti graminoids. African Journal of Ecology 23, 17994.
  • Crawley, M. J. (1983). Herbivory: the dynamics of animal-plant interactions. Blackwell Scientific Publications, Oxford.
  • Cristoffer, C. & Peres, C. A. (2003). Elephants versus butterflies: the ecological role of large herbivores in the evolutionary history of two tropical worlds. Journal of Biogeography 30, 13571380.
  • De Vries, W. (2009). Assessment of the relative importance of nitrogen deposition and climate change on the sequestration of carbon by forests in Europe: an overview introduction. Forest Ecology and Management 258(S1), vii–x.
  • De Vries, W., Reinds, G. J., Gundersen, P. & Sterba, H. (2006). The impact of nitrogen deposition on carbon sequestration in European forests and forest soils. Global Change Biology 12, 11511173.
  • Denman, K. L., Brasseur, G., Chidthaisong, A., Ciais, P., Cox, P. M., Dickinson, R. E., Hauglustaine, D., Heinze, C., Holland, E., Jacob, D., Lohmann, U., Ramachandran, S., Da Silva Dias, P. L., Wofsy, S. C. & Zhang, X. (2007). Couplings between changes in the climate system and biogeochemistry. In Climate change 2007: the physical science basis. Contribution of Working Group I to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change. (eds S. Solomon, D. Qin, M. Manning, Z. Chen, M. Marquis, K. B. Averyt, M. Tignor & H. L. Miller), pp. 499588. Cambridge University Press, Cambridge.
  • Derner, J., Boutton, T. & Briske, D. (2006). Grazing and ecosystem carbon storage in the North American Great Plains. Plant and Soil 280, 7790.
  • Doescher, P. S., Svejcar, T. J. & Jaindl, R. G. (1997). Gas exchange of Idaho fescue in response to defoliation and grazing history. Journal of Range Management 50, 285289.
  • Dolman, P. M. & Wäber, K. (2008). Ecosystem and competition impacts of introduced deer. Wildlife Research 35, 202214.
  • Dumont, B., Garel, J. P., Ginane, C., Decuq, F., Farruggia, A., Pradel, P., Rigolot, C. & Petit, M. (2007). Effect of cattle grazing a species-rich mountain pasture under different stocking rates on the dynamics of diet selection and sward structure. Animal 1, 10421052.
  • Dyer, M. I. (1980). Mammalian epidermal growth-factor promotes plant growth. Proceedings of the National Academy of Sciences 77, 48364837.
  • Ellis, E. C. & Ramankutty, N. (2008). Putting people in the map: anthropogenic biomes of the world. Frontiers in Ecology and the Environment 6, 439447.
  • Fearnside, P. M. (1995). Potential impacts of climatic change on natural forests and forestry in Brazilian Amazonia. Forest Ecology and Management 78, 5170.
  • Feeley, K. J. & Terborgh, J. W. (2005). The effects of herbivore density on soil nutrients and tree growth in tropical forest fragments. Ecology 86, 116124.
  • Feldpausch, T. R., Rondon, M. A., Fernandes, E. C. M., Riha, S. J. & Wandelli, E. (2004). Carbon and nutrient accumulation in secondary forests regenerating on pastures in central Amazonia. Ecological Applications 14, S164S176.
  • Food and Agriculture Organization of the United Nations [FAO]. (2000). Terrastat: human-induced land degradation due to agricultural activities. Food and Agriculture Organization of the United Nations, Rome.
  • Food and Agriculture Organization of the United Nations [FAO]. (2001). Global forest resources assessment 2000. Food and Agriculture Organization of the United Nations, Rome.
  • Food and Agriculture Organization of the United Nations [FAO]. (2008). FAOSTAT. Food and Agriculture Organization of the United Nations, Rome. http://faostat.fao.org/
  • Forbes, M. S., Raison, R. J. & Skjemstad, J. O. (2006). Formation, transformation and transport of black carbon (charcoal) in terrestrial and aquatic ecosystems. Science of the Total Environment 370, 190206.
  • Fornara, D. A. & Du Toit, J. (2008). Browsing-induced effects on leaf litter quality and decomposition in a southern African savanna. Ecosystems 11, 238249.
  • Forster, P., Ramaswamy, V., Artaxo, P., Berntsen, T., Betts, R., Fahey, D. W., Haywood, J., Lean, J., Lowe, D. C., Myhre, G., Nganga, J., Prinn, R., Raga, G., Schulz, M. & Van Dorland, R. (2007). Changes in atmospheric constituents and in radiative forcing. In Climate change 2007: the physical science basis. Contribution of Working Group I to the Fourth Assessment Report of the Intergovernmental Panel on Climate Change. (eds S. Solomon, D. Qin, M. Manning, Z. Chen, M. Marquis, K. B. Averyt, M. Tignor & H. L. Miller), pp. 129234. Cambridge University Press, Cambridge.
  • Forsyth, D. M. & Caley, P. (2006). Testing the irruptive paradigm of large-herbivore dynamics. Ecology 87, 297303.
  • Forsyth, D. M. & Duncan, R. P. (2001). Propagule size and the relative success of exotic ungulate and bird introductions to New Zealand. American Naturalist 157, 583595.
  • Frank, D. A. (1998). Ungulate regulation of ecosystem processes in Yellowstone National Park: direct and feedback effects. Wildlife Society Bulletin 26, 410418.
  • Frank, D. A. & Groffman, P. M. (1998). Ungulate versus landscape control of soil C and N processes in grasslands of Yellowstone National Park. Ecology 79, 22292241.
  • Frank, D. A., Kuns, M. M. & Guido, D. R. (2002). Consumer control of grassland plant production. Ecology 83, 602606.
  • Frank, D. A. & McNaughton, S. J. (1993). Evidence for the promotion of aboveground grassland production by native large herbivores in Yellowstone National Park. Oecologia 96, 157161.
  • Frank, A. B., Tanaka, D. L., Hofmann, L. & Follett, R. F. (1995). Soil carbon and nitrogen of Northern Great Plains grasslands as influenced by long-term grazing. Journal of Range Management 48, 470474.
  • Frelich, L. E. & Lorimer, G. G. (1985). Current and predicted long-term effects of deer browsing in hemlock forests in Michigan, USA. Biological Conservation 34, 99120.
  • Friedel, M. H. (1991). Range condition assessment and the concept of thresholds a viewpoint. Journal of Range Management 44, 422426.
  • Fritz, H. & Loison, A. (2006). Large herbivores across biomes. In Large herbivore ecology ecosystem dynamics and conservation. (eds K. Danell, R. Bergström, P. Duncan & J. Pastor), pp. 1949. Cambridge University Press, Cambridge.
  • Frost, C. J. & Hunter, M. D. (2004). Insect canopy herbivory and frass deposition affect soil nutrient dynamics and export in oak mesocosms. Ecology 85, 33353347.
  • Garel, M., Loison, A., Gaillard, J., Cugnasse, J. & Maillard, D. (2004). The effects of a severe drought on mouflon lamb survival. Proceedings of the Royal Society of London Series B-Biological Sciences 271, S471S473.
  • Gill, R. A. (2007). Influence of 90 years of protection from grazing on plant and soil processes in the subalpine of the Wasatch Plateau, USA. Rangeland Ecology and Management 60, 8898.
  • Gordon, I. J., Hester, A. J. & Festa-Bianchet, M. (2004). The management of wild large herbivores to meet economic, conservation and environmental objectives. Journal of Applied Ecology 41, 10211031.
  • Gornall, J., Jónsdóttir, I., Woodin, S. & Van Der Wal, R. (2007). Arctic mosses govern below-ground environment and ecosystem processes. Oecologia 153, 931941.
  • Grace, J. (2004). Presidential address: understanding and managing the global carbon cycle. Journal of Ecology 92, 189202.
  • Graz, F. P. (2008). The woody weed encroachment puzzle: gathering pieces. Ecohydrology 1, 340348.
  • Green, R. E., Cornell, S. J., Scharlemann, J. P. W. & Balmford, A. (2005). Farming and the fate of wild nature. Science 307, 550555.
  • Guldemond, R. & Van Aarde, R. (2008). A meta-analysis of the impact of African elephants on savanna vegetation. Journal of Wildlife Management 72, 892899.
  • Guo, L. B. & Gifford, R. M. (2002). Soil carbon stocks and land use change: a meta analysis. Global Change Biology 8, 345360.
  • Hamilton, E. W. & Frank, D. A. (2001). Can plants stimulate soil microbes and their own nutrient supply? Evidence from a grazing tolerant grass, Ecology 82, 23972402.
  • Hanley, N., Davies, A., Angelopoulos, K., Hamilton, A., Ross, A., Tinch, D. & Watson, F. (2008). Economic determinants of biodiversity change over a 400-year period in the Scottish uplands. Journal of Applied Ecology 45, 15571565.
  • Haukioja, E. & Koricheva, J. (2000). Tolerance to herbivory in woody vs. herbivorous plants. Evolutionary Ecology 14, 551562.
  • Hawkes, C. V. & Sullivan, J. J. (2001). The impact of herbivory on plants in different resource conditions: a meta-analysis. Ecology 82, 20452058.
  • Herms, D. A. & Mattson, W. J. (1992). The dilemma of plants—to grow or defend. Quarterly Review of Biology 67, 283335.
  • Hill, M., Roxburgh, S., Carter, J. & McKeon, G. (2005). Vegetation state change and consequent carbon dynamics in savanna woodlands of Australia in response to grazing, drought and fire: a scenario approach using 113 years of synthetic annual fire and grassland growth. Australian Journal of Botany 53, 715739.
  • Hirota, M., Tang, Y., Hu, Q., Kato, T., Hirata, S., Mo, W., Cao, G. & Mariko, S. (2005). The potential importance of grazing to the fluxes of carbon dioxide and methane in an alpine wetland on the Qinghai-Tibetan Plateau. Atmospheric Environment 39, 52555259.
  • Hobbs, N. T. (1996). Modification of ecosystems by ungulates. Journal of Wildlife Management 60, 695713.
  • Hobbs, R. J. & Norton, D. A. (1996). Towards a conceptual framework for restoration ecology. Restoration Ecology 4, 93110.
  • Höft, R. & Höft, M. (1995). The differential effects of elephants on rain forest communities in the Shimba Hills, Kenya. Biological Conservation 73, 6779.
  • Holdo, R. M., Sinclair, A. R. E., Dobson, A. P., Metzger, K. L., Bolker, B. M., Ritchie, M. E. & Holt, R. D. (2009). A disease-mediated trophic cascade in the Serengeti and its implications for ecosystem C. PLoS Biology 7, e1000210.
  • Holland, E. A., Parton, W. J., Detling, J. K. & Coppock, D. L. (1992). Physiological responses of plant populations to herbivory and their consequences for ecosystem nutrient flow. American Naturalist 140, 685706.
  • Holland, J. N., Cheng, W. & Crossley, D. A. (1996). Herbivore-induced changes in plant carbon allocation: assessment of below-ground C fluxes using carbon-14. Oecologia 107, 8794.
  • Holland, J. P., Waterhouse, A., Robertson, D. & Pollock, M. L. (2008). Effect of different grazing management systems on the herbage mass and pasture height of a Nardus stricta grassland in western Scotland, United Kingdom. Grass and Forage Science 63, 4859.
  • Holling, C. S. (1973). Resilience and stability of ecological systems. Annual Review of Ecology and Systematics 4, 123.
  • Horner, J., Gosz, J. & Cates, R. (1988). The role of carbon-based plant secondary metabolites in decomposition in terrestrial ecosystems. American Naturalist 132, 869883.
  • Houghton, R. A. (2003). Revised estimates of the annual net flux of carbon to the atmosphere from changes in land use 1850-2000. Tellus 55B, 378–390.
  • Houghton, R. A. (2007). Balancing the global carbon budget. Annual of Review Earth Planetary Sciences 35, 31347.
  • Houghton, R. A. & Goodale, C. L. (2004). Effects of land-use change on the carbon balance of terrestrial ecosystems. In Ecosystems and land use change (eds R. DeFries, G. Asner & R. A. Houghton), pp. 8598. American Geophysical Union, Washington, DC.
  • Houghton, R. A., Hackler, J. L. & Lawrence, K. T. (1999). The US carbon budget: contributions from land-use change. Science 285, 574578.
  • Houghton, R. A., Hobbie, J. E., Melillo, J. M., More, B., Peterson, B. J., Shaver, G. R. & Woodwell, G. M. (1983). Changes in the carbon cycle in terrestrial biota and soils between 1860 and 1980: a net release of CO2 to the atmosphere. Ecological Monographs 53, 235262.
  • Howe, H. F., Zorn-Arnold, B., Sullivan, A. & Brown, J. S. (2006). Massive and distinctive effects of meadow voles on grassland vegetation. Ecology 87, 30073013.
  • Hungate, B. A., Holland, E. A., Jackson, R. B., Chapin, F. S., Mooney, H. A. & Field, C. B. (1997). The fate of carbon in grasslands under carbon dioxide enrichment. Nature 388, 576579.
  • Huntly, N. (1991). Herbivores and the dynamics of communities and ecosystems. Annual Review of Ecology and Systematics 22, 477503.
  • Huston, M. A. & Wolverton, S. (2009). The global distribution of net primary production: resolving the paradox. Ecological Monographs 79, 343377.
  • Ickes, K. (2001). Hyper-abundance of native wild pigs (Sus scrofa) in a lowland dipterocarp rain forest of peninsular Malaysia. Biotropica 33, 682690.
  • Ickes, K., Dewalt, S. & Appanah, S. (2001). Effects of native pigs (Sus scrofa) on woody understorey vegetation in a Malaysian lowland rain forest. Journal of Tropical Ecology 17, 191206.
  • Ise, T., Dunn, A. L., Wofsy, S. C. & Moorcroft, P. R. (2008). High sensitivity of peat decomposition to climate change through water-table feedback. Nature Geoscience 1, 763766.
  • Jackson, R. B., Banner, J. L., Jobbágy, E. G., Pockman, W. T. & Wall, D. H. (2002). Ecosystem carbon loss with woody plant invasion of grasslands. Nature 418, 623626.
  • Jacobs, S. & Naiman, R. (2008). Large African herbivores decrease herbaceous plant biomass while increasing plant species richness in a semi-arid savanna toposequence. Journal of Arid Environments 72, 891903.
  • Jandl, R., Lindner, M., Vesterdal, L., Bauwens, B., Baritz, R., Hagedorn, F., Johnson, D. W., Minkkinen, K. & Byrne, K. A. (2007). How strongly can forest management influence soil carbon sequestration? Geoderma 137, 253268.
  • Jaramillo, V. & Detling, J. K. (1988). Grazing history, defoliation, and competition: effects on shortgrass production and nitrogen accumulation. Ecology 69, 15991608.
  • Jauregui, B. M., Rosa-Garcia, R., Garcia, U., Wallisdevries, M. F., Osoro, K. & Celaya, R. (2008). Effects of stocking density and breed of goats on vegetation and grasshopper occurrence in heathlands. Agriculture, Ecosystems & Environment 123, 219224.
  • Jobbágy, E. G. & Jackson, R. B. (2000). The vertical distribution of soil organic carbon and its relation to climate and vegetation. Ecological Applications 10, 423436.
  • Johnston, K. M. & Schmitz, O. J. (1997). Wildlife and climate change: assessing the sensitivity of selected species to simulated doubling of atmospheric CO2. Global Change Biology 3, 531544.
  • Jonard, M., Misson, L. & Ponette, Q. (2006). Long-term thinning effects on the forest floor and the foliar nutrient status of Norway spruce stands in the Belgian Ardennes. Canadian Journal of Forest Research 36, 26842695.
  • Junk, W., Brown, M., Campbell, I., Finlayson, M., Gopal, B., Ramberg, L. & Warner, B. (2006). The comparative biodiversity of seven globally important wetlands: a synthesis. Aquatic Sciences 68, 400414.
  • Kaimowitz, D., Mertens, B., Wunder, S. & Pacheco, P. (2004). Hamburger connection fuels Amazon destruction. Centre for International Forestry Research, Bogor.
  • Kapos, V., Ravilious, C., Campbell, A., Dickson, B., Gibbs, H., Hansen, M., Lysenko, I., Miles, L., Price, J., Scharlemann, J. P. W. & Trumper, K. (2008). Carbon and biodiversity: a demonstration atlas. UNEP-WCMC, Cambridge.
  • Keith, H., Mackey, B. G. & Lindenmayer, D. B. (2009). Re-evaluation of forest biomass carbon stocks and lessons from the world's most carbon-dense forests. Proceedings of the National Academy of Sciences 106, 1163511640.
  • Keesing, F. (2000). Cryptic consumers and the ecology of an African savanna. BioScience 50, 205215.
  • Kieft, T. L. (1994). Grazing and plant-canopy effects on semiarid soil microbial biomass and respiration. Biology and Fertility of Soils 18, 155162.
  • Kielland, K. & Bryant, J. P. (1998). Moose herbivory in taiga: effects on biogeochemistry and vegetation dynamics in primary succession. Oikos 82, 377383.
  • Knapp, A. K., Blair, J. M., Briggs, J. M., Collins, S. L., Hartnett, D. C., Johnson, L. C. & Towne, E. G. (1999). The keystone role of bison in North American tallgrass prairie. BioScience 49, 3950.
  • Koh, S. (2002). The response of four species of spring flowering perennial herbs to grazing by white-tailed deer in southern Ontario: a multiple-scale evaluation. PhD Thesis, York University, Toronto.
  • Kruska, R. L., Reid, R. S., Thornton, P. K., Henninger, N. & Kristjanson, P. M. (2003). Mapping livestock-oriented agricultural production systems for the developing world. Agricultural Systems 77, 3963.
  • Kunhamu, T., Kumar, B. & Viswanath, S. (2009). Does thinning affect litterfall, litter decomposition, and associated nutrient release in Acacia mangium stands of Kerala in peninsular India? Canadian Journal of Forest Research 39, 792801.
  • Lal, R. (2002). Carbon sequestration in dryland ecosystems of West Asia and North Africa. Land Degradation and Development 13, 4559.
  • Lal, R. (2003). Offsetting global CO2 emissions by restoration of degraded soils and intensification of world agriculture and forestry. Land Degradation and Development 14, 309322.
  • Lal, R. (2004). Carbon sequestration in dryland ecosystems. Environmental Management 33, 528544.
  • Lawes, M. & Chapman, C. (2006). Does the herb Acanthus pubescens and/or elephants suppress tree regeneration in disturbed Afrotropical forest? Forest Ecology and Management 221, 278284.
  • Laycock, W. A. (1991). Stable states and thresholds of range condition on North American rangelands: a viewpoint. Journal of Range Management 44, 42733.
  • Lecain, D. R., Morgan, J. A., Schuman, G. E., Reeder, J. D. & Hart, R. H. (2002). Carbon exchange and species composition of grazed pastures and exclosures in the shortgrass steppe of Colorado. Agriculture, Ecosystems & Environment 93, 421435.
  • Lehmann, J., Gaunt, J. & Rondon, M. (2006). Bio-char sequestration in terrestrial ecosystems: a review. Mitigation and Adaptation Strategies for Global Change 11, 403427.
  • Lehner, B. & Döll, P. (2004). Development and validation of a global database of lakes, reservoirs and wetlands. Journal of Hydrology 296, 122.
  • Leopold, A., Sowls, L. K. & Spencer, D. L. (1947). A survey of over-populated deer ranges in the United States. Journal of Wildlife Management 11, 162177.
  • Leriche, H., Leroux, X., Gignoux, J., Tuzet, A., Fritz, H., Abbadie, L. & Loreau, M. (2001). Which functional processes control the short-term effect of grazing on net primary production in grasslands? Oecologia 129, 114124.
  • Levinsky, I., Skov, F., Svenning, J. & Rahbek, C. (2007). Potential impacts of climate change on the distributions and diversity patterns of European mammals. Biodiversity and Conservation 16, 38033816.
  • Lewis, S. L., Lopez-Gonzalez, G., Sonke, B., Affum-Baffoe, K., Baker, T. R., Ojo, L. O., Phillips, O. L., Reitsma, J. M., White, L., Comiskey, J. A., Djuikouo, M. N., Ewango, C. E. N., Feldpausch, T. R., Hamilton, A. C., Gloor, M., Hart, T., Hladik, A., Lloyd, J., Lovett, J. C., Makana, J., Malhi, Y., Mbago, F. M., Ndangalasi, H. J., Peacock, J., Peh, K. S., Sheil, D., Sunderland, T., Swaine, M. D., Taplin, J., Taylor, D., Thomas, S. C., Votere, R. & Woll, H. (2009). Increasing carbon storage in intact African tropical forests. Nature 457, 10031006.
  • Li, S., Asanuma, J., Eugster, W., Kotani, A., Liu, J. J., Urano, T., Oikawa, T., Davaa, G., Oyunbaatar, D. & Sugita, M. (2005). Net ecosystem carbon dioxide exchange over grazed steppe in central Mongolia. Global Change Biology 11, 19411955.
  • Litton, C. M., Raich, J. W. & Ryan, M. G. (2007). Carbon allocation in forest ecosystems. Global Change Biology 13, 20892109.
  • Loison, A., Jullien, J. & Menaut, P. (1999). Relationship between chamois and isard survival and variation in global and local climate regimes: contrasting examples from the Alps and Pyrenees. Ecological Bulletins 47, 126136.
  • Lovett, G. M., Canham, C. D., Arthur, M. A., Weathers, K. C. & Fitzhugh, R. D. (2006). Forest ecosystem responses to exotic pests and pathogens in eastern North America. BioScience 56, 395.
  • Ludwig, J. A. & Tongway, D. J. (1995). Desertification in Australia: an eye to grass roots and landscapes. Environmental Monitoring and Assessment 37, 321327.
  • Luyssaert, S., Schulze, E., Borner, A., Knohl, A., Hessenmoller, D., Law, B. E., Ciais, P. & Grace, J. (2008). Old-growth forests as global carbon sinks. Nature 455, 213215.
  • Mack, M. C., Schuur, E. A. G., Bret-Harte, M. S., Shaver, G. R. & Chapin, F. S. (2004). Ecosystem carbon storage in arctic tundra reduced by long-term nutrient fertilization. Nature 431, 440443.
  • Magnani, F., Mencuccini, M., Borghetti, M., Berbigier, P., Berninger, F., Delzon, S., Grelle, A., Hari, P., Jarvis, P. G., Kolari, P., Kowalski, A. S., Lankreijer, H., Law, B. E., Lindroth, A., Loustau, D., Manca, G., Moncrieff, J. B., Rayment, M., Tedeschi, V., Valentini, R. & Grace, J. (2007). The human footprint in the carbon cycle of temperate and boreal forests. Nature 447, 849851.
  • Maronna, R. A., Martin, D. J. & Yohai, V. J. (2006). Robust statistics: theory and methods. Wiley, Chichester.
  • Martin, P. S. (1984). Prehistoric overkill: the global model. In Quaternary extinctions: a prehistoric revolution. (eds P. S. Martin & R. G. Klein), pp. 354–403. University of Arizona Press, Tuscon.
  • Matthews, E. (1997). Global litter production, pools, and turnover times: Estimates from measurement data and regression models. Journal of Geophysical Research 102, 18771800.
  • May, R. M. (1977). Thresholds and breakpoints in ecosystems with a multiplicity of stable states. Nature 269, 471477.
  • McGuire, A. D., Sitch, S., Clein, J. S., Dargaville, R., Esser, G., Foley, J., Heimann, M., Joos, F., Kaplan, J., Kicklighter, D. W., Meier, R. A., Melillo, J. M., Moore, B., Prentice, I. C., Ramankutty, N., Reichenau, T., Schloss, A., Tian, H., Williams, L. J. & Wittenberg, U. (2001). Carbon balance of the terrestrial biosphere in the twentieth century: analyses of CO2, climate and land-use effects with four process-based ecosystem models. Global Biogeochemical Cycles 15, 183206.
  • McInnes, P. F., Naiman, R. J., Pastor, J. & Cohen, Y. (1992). Effects of moose browsing on vegetation and litter of the boreal forest, Isle Royale, Michigan, USA. Ecology 73, 20592075.
  • McMahon, C. & Bradshaw, C. (2008). To catch a buffalo: field immobilisation of Asian swamp buffalo using etorphine and xylazine. Australian Veterinary Journal 86, 235241.
  • McMahon, S. M., Parker, G. G. & Miller, D. R. (2010). Evidence for a recent increase in forest growth. Proceedings of the National Academy of Sciences 107, 36113615.
  • McNaughton, S. J. (1984). Grazing lawns: animals in herds, plant form, and coevolution. American Naturalist 124, 863886.
  • McNaughton, S. J. (1985). Ecology of a grazing system: the Serengeti. Ecological Monographs 55, 259294.
  • Mcnaughton, S. J. & Georgiadis, N. J. (1986). Ecology of African grazing and browsing mammals. Annual Review of Ecology and Systematics 17, 3965.
  • Mcnaughton, S. J., Oesterheld, M., Frank, D. A. & Williams, K. J. (1989). Ecosystem-level patterns of primary productivity and herbivory in terrestrial habitats. Nature 341, 142144.
  • McNaughton, S. J., Banyikwa, F. F. & McNaughton, M. M. (1997). Promotion of the cycling of diet-enhancing nutrients by African grazers. Science 278, 17981800.
  • Meeuwig, R. O. (1965). Effects of seeding and grazing on infiltration capacity and soil stability of a subalpine range in central Utah. Journal of Range Management 18, 173180.
  • Milchunas, D. G. & Lauenroth, W. K. (1993). Quantitative effects of grazing on vegetation and soils over a global range of environments. Ecological Monographs 63, 327366.
  • Miller, G. H., Fogel, M. L., Magee, J. W., Gagan, M. K., Clarke, S. J. & Johnson, B. J. (2005). Ecosystem collapse in pleistocene Australia and a human role in megafaunal extinction. Science 309, 287290.
  • Mills, A. J. & Cowling, R. M. (2006). Rate of carbon sequestration at two thicket restoration sites in the eastern Cape, South Africa. Restoration Ecology 14, 3849.
  • Miyaki, M. & Kaji, K. (2009). The dynamics of forest stands affected by sika deer on Nakanoshima Island—Change of size structure similar to the thinning effect. In Sika Deer: biology and management of native and introduced populations. (eds D. R. McCullough, S. Takatsuki & K. Kaji), pp. 181191. Springer, Tokyo.
  • Morris, J. T. & Jensen, A. (1998). The carbon balance of grazed and non-grazed Spartina anglica saltmarshes at Skallingen, Denmark. Journal of Ecology 86, 229242.
  • Mouillot, F., Narasimha, A., Balkanski, Y., Lamarque, J. & Field, C. B. (2006). Global carbon emissions from biomass burning in the 20th century. Geophysical Research Letters 33, L01801.
  • Mountford, E. P. & Peterken, G. E. (2003). Long-term change and implications for the management of wood-pastures: experience over 40 years from Denny Wood, New Forest. Forestry 76, 1943.
  • Nadelhoffer, K. J., Emmett, B. A. & Gundersen, P. (1999). Nitrogen deposition makes a minor contribution to carbon sequestration in temperate forests. Nature 398, 145148.
  • Neff, J. C., Townsend, A. R., Gleixner, G., Lehman, S. J., Turnbull, J. & Bowman, W. D. (2002). Variable effects of nitrogen additions on the stability and turnover of soil carbon. Nature 419, 915917.
  • Nepstad, D., Soares-Filho, B., Merry, F., Moutinho, P., Rodrigues, H. O., Bowman, M., Schwartzman, S., Almeida, O. & Rivero, S. (2007). The costs and benefits of reducing carbon emissions from deforestation and forest degradation in the Brazilian Amazon. Woods Hole Research Center, Falmouth.
  • Niemelä, P., Chapin, F., Danell, K. & Bryant, J. (2001). Herbivory mediated responses of selected boreal forests to climatic change. Climatic Change 48, 427440.
  • Nomiya, H., Suzuki, W., Kanzashi, T., Shibata, M., Tanaka, H. & Nakashizuka, T. (2002). The response of forest floor vegetation and tree regeneration to deer exclusion and disturbance in a riparian deciduous forest, central Japan. Plant Ecology 164, 263276.
  • Nowacki, G. J. & Abrams, M. D. (2008). The demise of fire and “mesophication” of forests in the eastern United States. BioScience 58, 123138.
  • Nowak, R. M. (1999). Walker's mammals of the world. John Hopkins University Press, Baltimore.
  • Ogutu, J. & Owen-Smith, N. (2003). ENSO, rainfall and temperature influences on extreme population declines among African savanna ungulates. Ecology Letters 6, 412419.
  • Olff, H., Ritchie, M. E. & Prins, H. H. T. (2002). Global environmental controls of diversity in large herbivores. Nature 415, 901904.
  • Olofsson, J., Kitti, H., Rautiainen, P., Stark, S. & Oksanen, L. (2001). Effects of summer grazing by reindeer on composition of vegetation, productivity and nitrogen cycling. Ecography 24, 1324.
  • Olofsson, J. & Oksanen, L. (2002). Role of litter decomposition for the increased primary production in areas heavily grazed by reindeer: a litterbag experiment. Oikos 96, 507515.
  • Olofsson, J., Stark, S. & Oksanen, L. (2004). Reindeer influence on ecosystem processes in the tundra. Oikos 105, 38696.
  • Owen, D. F. & Wiegert, R. G. (1981). Mutualism between grasses and grazers: an evolutionary hypothesis. Oikos 36, 376378.
  • Owensby, C. E., Ham, J. M. & Auen, L. M. (2006). Fluxes of CO2 from grazed and ungrazed tallgrass prairie. Rangeland Ecology and Management 59, 111127.
  • Owen-Smith, N. (1987). Pleistocene extinctions: the pivotal role of megaherbivores. Paleobiology 13, 351362.
  • Pacala, S. W., Hurtt, G. C., Baker, D., Peylin, P., Houghton, R. A., Birdsey, R. A., Heath, L., Sundquist, E. T., Stallard, R. F., Ciais, P., Moorcroft, P., Caspersen, J. P., Shevliakova, E., Moore, B., Kohlmaier, G., Holland, E., Gloor, M., Harmon, M. E., Fan, S., Sarmiento, J. L., Goodale, C. L., Schimel, D. & Field, C. B. (2001). Consistent land- and atmosphere-based U.S. carbon sink estimates. Science 292, 23162320.
  • Paige, K. N. & Whitham, T. G. (1987). Overcompensation in response to mammalian herbivory: the advantage of being eaten. American Naturalist 129, 407416.
  • Paine, C. E. T. & Beck, H. (2007). Seed predation by Neotropical rainforest mammals increases diversity in seedling recruitment. Ecology 88, 30733087.
  • Palmer, T. M., Stanton, M. L., Young, T. P., Goheen, J. R., Pringle, R. M. & Karban, R. (2008). Breakdown of an ant-plant mutualism follows the loss of large herbivores from an African savanna. Science 319, 192195.
  • Parish, F., Sirin, A., Charman, D., Joosten, H., Minayeva, T., Silvius, M., Stringer, L. (eds) (2008). Assessment on peatlands, biodiversity and climate change: main report. Global Environment Centre and Wetlands International, Kuala Lumpur.
  • Pastor, J., Dewey, B., Naiman, R. J., Mcinnes, P. F. & Cohen, Y. (1993). Moose browsing and soil fertility in the boreal forests of Isle Royale National Park. Ecology 74, 467480.
  • Peltzer, D. A., Allen, R. B., Lovett, G. M., Whitehead, D. & Wardle, D. A. (2010). Effects of biological invasions on forest carbon sequestration. Global Change Biology 16, 732746.
  • Piñeiro, G., Paruelo, J. M., Jobbágy, E. G., Jackson, R. B. & Oesterheld, M. (2009). Grazing effects on belowground C and N stocks along a network of cattle exclosures in temperate and subtropical grasslands of South America. Global Biogeochemical Cycles 23, GB2003.
  • Piñeiro, G., Paruelo, J. M. & Oesterheld, M. (2006). Potential long-term impacts of livestock introduction on carbon and nitrogen cycling in grasslands of Southern South America. Global Change Biology 12, 12671284.
  • Ping, C., Michaelson, G. J., Jorgenson, M. T., Kimble, J. M., Epstein, H., Romanovsky, V. E. & Walker, D. A. (2008). High stocks of soil organic carbon in the North American Arctic region. Nature Geoscience 1, 615619.
  • Post, E. & Stenseth, N. (1999). Climatic variability, plant phenology, and northern ungulates. Ecology 80, 13221339.
  • Prins, H. T. & Van Der Jeugd, H. P. (1993). Herbivore population crashes and woodland structure in East Africa. Journal of Ecology 81, 305314.
  • Pucheta, E., Cabido, M., Díaz, S. & Funes, G. (1998). Floristic composition, biomass and aboveground net plant production in grazed and protected sites in a mountain grassland of central Argentina. Acta Oecologica 19, 97105.
  • R Development Core Team. (2008). R: a language and environment for statistical computing. R Foundation for Statistical Computing, Vienna.
  • Ramankutty, N., Evan, A. T., Monfred, A. C. & Foley, J. A. (2008). Farming the planet: 1. Geographic distribution of global agricultural lands in the year 2000. Global Biogeochemical Cycles 22, GB1003.
  • Rangel-Castro, J. I., Prosser, J. I., Scrimgeour, C. M., Smith, P., Ostle, N., Ineson, P., Meharg, A. & Killham, K. (2004). Carbon flow in an upland grassland: effect of liming on the flux of recently photosynthesized carbon to rhizosphere soil. Global Change Biology 10, 21002108.
  • Rautiainen, A., Saikku, L. & Kauppi, P. E. (2010). Carbon gains and recovery from degradation of forest biomass in European Union during 1990–2005. Forest Ecology and Management 259, 12321238.
  • Reeder, J. D. & Schuman, G. E. (2002). Influence of livestock grazing on C sequestration in semi-arid mixed grass and short-grass rangelands. Environmental Pollution 116, 457463.
  • Reid, R., Galvin, K. & Kruska, R. (2008). Global significance of extensive grazing lands and pastoral societies: an introduction. In Fragmentation in semi-arid and arid landscapes. (eds K. A. Galvin, R. S. Reid, R. H. Behnke & N. T. Hobbs), pp. 124. Springer, Dordrecht.
  • Rhemtulla, J. M., Mladenoff, D. J. & Clayton, M. K. (2009). Historical forest baselines reveal potential for continued carbon sequestration. Proceedings of the National Academy of Sciences 106, 60826087.
  • Rietkerk, M., Ketner, P., Stroosnijder, L. & Prins, H. H. T. (1996). Sahelian rangeland development: a catastrophe? Journal of Range Management 4, 512519.
  • Risch, A. C. & Frank, D. A. (2006). Carbon dioxide fluxes in a spatially and temporally heterogeneous temperate grassland. Oecologia 147, 291302.
  • Robinson, D. (2007). Implications of a large global root biomass for carbon sink estimates and for soil carbon dynamics. Proceedings of the Royal Society B 274, 27532759.
  • Rooke, T. (2003). Growth responses of a woody species to clipping and goat saliva. African Journal of Ecology 41, 324328.
  • Royo, A. A. & Carson, W. P. (2005). The herb community of a tropical forest in central Panama: dynamics and impact of mammalian herbivores. Oecologia 145, 6675.
  • Saikku, L., Rautiainen, A. & Kauppi, P. E. (2008). The sustainability challenge of meeting carbon dioxide targets in Europe by 2020. Energy Policy 36, 730742.
  • Sankaran, M. & Augustine, D. J. (2004). Large herbivores suppress decomposer abundance in a semiarid grazing ecosystem. Ecology 85, 10521061.
  • Sarmiento, G. (1992). Adaptive strategies of perennial grasses in South American savannas. Journal of Vegetation Science 3, 325336.
  • Saugier, B., Roy, J. & Mooney, H. A. (2001). Estimations of global terrestrial productivity: converging toward a single number? In Terrestrial global productivity. (eds J. Roy, B. Saugier & H. A. Mooney), pp. 543558. Academic Press, San Diego.
  • Schlesinger, W. H. & Lichter, J. (2001). Limited carbon storage in soil and litter of experimental forest plots under increased atmospheric CO2. Nature 411, 466469.
  • Schmitt, C. B., Burgess, N. D., Coad, L., Belokurov, A., Besançon, C., Boisrobert, L., Campbell, A., Fish, L., Gliddon, D., Humphries, K., Kapos, V., Loucks, C., Lysenko, I., Miles, L., Mills, C., Minnemeyer, S., Pistorius, T., Ravilious, C., Steininger, M. & Winkel, G. (2009). Global analysis of the protection status of the world's forests. Biological Conservation 142, 21222130.
  • Schnabel, R. R., Franzluebbers, A. J., Stout, W. L., Sanderson, M. A. & Stuedemann, J. A. (2001). The effects of pasture management practices. In The potential of U.S. grazing lands to sequester carbon and mitigate the greenhouse effect. (eds R. F. Follett, J. M. Kimble & R. Lal), pp. 291322. Lewis Publishers, Boca Raton.
  • Schuman, G. E., Reeder, J. D., Manley, J. T., Hart, R. H. & Manley, W. A. (1999). Impact of grazing management on the carbon and nitrogen balance of a mixed-grass rangeland. Ecological Applications 9, 6571.
  • Semmartin, M. & Ghersa, C. M. (2006). Intra-specific changes in plant morphology, associated with grazing, and effects on litter quality, carbon and nutrient dynamics during decomposition. Austral Ecology 31, 99105.
  • Shannon, G., Druce, D. J., Page, B. R., Eckhardt, H. C., Grant, R. & Slotow, R. (2008). The utilization of large savanna trees by elephant in southern Kruger National Park. Journal of Tropical Ecology 24, 281289.
  • Shipley, L. A., Gross, J. E., Spalinger, D. E., Hobbs, N. T. & Wunder, B. A. (1994). The scaling of intake rate in mammalian herbivores. American Naturalist 143, 10551082.
  • Shutou, K. & Nakane, K. (2004). Change in soil carbon cycling for stand development of Japanese cedar (Cryptomeria japonica) plantations following clear-cutting. Ecological Research 19, 233244.
  • Silvester, W. B. & Orchard, T. A. (1999). The biology of kauri (Agathis australis) in New Zealand. I. Production, biomass, carbon storage, and litter fall in four forest remnants. New Zealand Journal of Botany 37, 553571.
  • Sinclair, A. R E. (1979). The eruption of the ruminants. In Serengeti: dynamics of an ecosystem. (eds A. R. E. Sinclair & M. Norton-Griffiths), pp. 82103. University of Chicago Press, Chicago.
  • Sinclair, A. R. E. & Fryxell, J. M. (1985). The Sahel of Africa: ecology of a disaster. Canadian Journal of Zoology 63, 98794.
  • Singer, F. J. & Schoenecker, K. A. (2003). Do ungulates accelerate or decelerate nitrogen cycling? Forest Ecology and Management 181, 189204.
  • Sinton, P., Seastedt, T., Bennett, B., Bock, C. & Bock, J. (2000). C3 and C4 species changes identified by δ13C values of soil organic matter in a Colorado prairie. Western North American Naturalist 60, 445450.
  • Sjögersten, S., Van Der Wal, R. & Woodin, S. J. (2008). Habitat type determines herbivory controls over CO2 fluxes in a warmer Arctic. Ecology 89, 21032116.
  • Skinner, R. H. (2008). High biomass removal limits carbon sequestration potential of mature temperate pastures. Journal of Environmental Quality 37, 13191326.
  • Smit, R., Bokdam, J., Den Ouden, J., Olff, H., Schot-Opschoor, H. & Schrijvers, M. (2001). Effects of introduction and exclusion of large herbivores on small rodent communities. Plant Ecology 155, 119127.
  • Solbrig, O. T. (2006). Economic growth and environmental change. In The Cambridge economic history of Latin America: the long twentieth century. (eds V. Bulmer-Thomas, J. H. Coatsworth & R. C. Conde), pp. 329376. Cambridge University Press, Cambridge.
  • Soussana, J. F., Loiseau, P., Vuichar, N., Ceschia, E., Balesdent, J., Chevallier, T. & Arrouays, D. (2004). Carbon cycling and sequestration opportunities in temperate grasslands. Soil Use and Management 20, 219230.
  • Stark, S. & Grellmann, D. (2002). Soil microbial responses to herbivory in an Arctic tundra heath at two levels of nutrient availability. Ecology 83, 27362745.
  • Steen, H., Mysterud, A. & Austrheim, G. (2005). Sheep grazing and rodent populations: evidence of negative interactions from a landscape scale experiment. Oecologia 143, 357364.
  • Steinfeld, H., Gerber, P., Wassenaar, T., Castel, V., Rosales, M. & De Haan, C. (2006). Livestock's long shadow: environmental issues and options. Food and Agriculture Organization of the United Nations, Rome.
  • Stock, W. D., Roux, D. & Heyden, F. (1993). Regrowth and tannin production in woody and succulent karoo shrubs in response to simulated browsing. Oecologia 96, 562568.
  • Strassmann, K., Joos, F. & Fischer, G. (2008). Simulating effects of land use changes on carbon fluxes: past contributions to atmospheric CO2 increases and future commitments due to losses of terrestrial sink capacity. Tellus B 60, 583603.
  • Susiluoto, S., Rasilo, T., Pumpanen, J. & Berninger, F. (2008). Effects of grazing on the vegetation structure and carbon dioxide exchange of a Fennoscandian fell ecosystem. Arctic, Antarctic, and Alpine Research 40, 422431.
  • Takatsuki, S. (2009). Effects of sika deer on vegetation in Japan: a review. Biological Conservation 142, 19221929.
  • Thomas, R. Q., Canham, C. D., Weathers, K. C. & Goodale, C. L. (2009). Increased tree carbon storage in response to nitrogen deposition in the US. Nature Geoscience 3, 1317.
  • Thuiller, W., Broennimann, O., Hughes, G., Alkemade, J. R. M., Midgley, G. F. & Corsi, F. (2006). Vulnerability of African mammals to anthropogenic climate change under conservative land transformation assumptions. Global Change Biology 12, 424440.
  • Tolvanen, A. & Laine, K. (1997). Effects of reproduction and artificial herbivory on vegetative growth and resource levels in deciduous and evergreen dwarf shrubs. Canadian Journal of Botany 75, 656666.
  • Towne, E. G., Hartnett, D. C. & Cochran, R. C. (2005). Vegetation trends in tallgrass prairie from bison and cattle grazing. Ecological Applications 15, 15501559.
  • Tremblay, J. P., Huot, J. & Potvin, F. (2006). Divergent nonlinear responses of the boreal forest field layer along an experimental gradient of deer densities. Oecologia 150, 7888.
  • Trumbore, S. (2006). Carbon respired by terrestrial ecosystems—recent progress and challenges. Global Change Biology 12, 141153.
  • Ung, C. H., Bernier, P. & Guo, X. J. (2008). Canadian national biomass equations: new parameter estimates that include British Columbia data. Canadian Journal of Forest Research 38, 11231132.
  • Van Der Wal, R. (2006). Do herbivores cause habitat degradation or vegetation state transition? Evidence from the tundra. Oikos 114, 177186.
  • Van Wieren, S. E. (1995). The potential role of large herbivores in nature conservation and extensive land use in Europe. Biological Journal of the Linnean Society 56(S1), 1123.
  • Vera, F. W. M. (2002). Grazing ecology and forest history. CABI Publishing, Wallingford.
  • Virtanen, R., Salminen, J. & Strömmer, R. (2008). Soil and decomposer responses to grazing exclusion are weak in mountain snow-beds. Acta Oecologica 33, 207212.
  • Vors, L. S. & Boyce, M. S. (2009). Global declines of caribou and reindeer. Global Change Biology 15, 26262633.
  • Waldram, M. S., Bond, W. J. & Stock, W. D. (2008). Ecological engineering by a mega-grazer: white rhino impacts on a South African savanna. Ecosystems 11, 101112.
  • Ward, S. E., Bardgett, R. D., McNamara, N. P., Adamson, J. K & Ostle, N. J. (2007). Long-term consequences of grazing and burning on northern peatland carbon dynamics. Ecosystems 10, 10691083.
  • Wardle, D. A., Barker, M. B., Yeates, G. W., Bonner, K. I. & Ghani, A. (2001). Introduced browsing mammals in New Zealand natural forests: aboveground and belowground consequences. Ecological Monographs 71, 587614.
  • Wardle, D. A., Bonner, K. I. & Baker, G. M. (2002). Linkages between plant litter decomposition, litter quality, and vegetation responses to herbivores. Functional Ecology 16, 585595.
  • Watt, A. S. (1947). Pattern and process in the plant community. Journal of Ecology 35, 122.
  • Webb, S. D. (1978). A history of savanna vertebrates in the New World. Part II: South America and the Great Interchange. Annual Review of Ecology and Systematics 9, 393426.
  • Weber, P., Rigling, A. & Bugmann, H. (2008). Investigating the impact of grazing on the dynamics of mixed Pinus sylvestris and Quercus pubescens stands by applying a forest gap model. Ecological Modelling 210, 301311.
  • Weinhold, B. J., Hendrickson, J. R. & Karn, J. F. (2001). Pasture management influences on soil properties in the Northern Great Plains. Journal of Soil and Water Conservation 56, 2731.
  • Welker, J. M., Fahnestock, J. T., Povirk, K. L., Bilbrough, C. J. & Piper, R. E. (2004). Alpine grassland CO2 exchange and nitrogen cycling: grazing history effects, Medicine Bow Range, Wyoming, USA. Arctic Antarctic and Alpine Research 36, 1120.
  • Westoby, M., Walker, B. & Noy-Meir, I. (1989). Opportunistic management for rangelands not at equilibrium. Journal of Range Management 42, 26674.
  • White, R. P., Murray, S. & Rohweder, M. (2000). Pilot analysis of global ecosystems: grassland ecosystems. World Resources Institute, Washington, DC.
  • Wilsey, B. J., Parent, G., Roulet, N. T., Moore, T. R. & Potvin, C. (2002). Tropical pasture carbon cycling: relationships between C source/sink strength, above-ground biomass and grazing. Ecology Letters 5, 367376.
  • Wissenschaftlicher Beirat der Bundesregierung Globale Umweltveränerungen [WBGU]. (1988). Die Anrechnung biolischer Quellen und Senken in Kyoto-Protokoll: Fortschritt oder Rückschlang für den globalen Umweltschutz Sondergutachten 1988. Wissenschaftlicher Beirat der Bundesregierung Globale Umweltveränerungen, Bremerhaven.
  • Wolf, E. C., Cooper, D. J. & Hobbs, N. T. (2007). Hydrologic regime and herbivory stabilize an alternative state in Yellowstone National Park. Ecological Applications 17, 15721587.
  • Wood, M. K. & Blackburn, W. H. (1981). Grazing systems: their influence on infiltration rates in the rolling plains of Texas. Journal of Range Management 34, 331335.
  • Woodbury, P. B., Heath, L. S. & Smith, J. E. (2006). Land use change effects on forest carbon cycling throughout the southern United States. Journal of Environmental Quality 35, 13481363.
  • Wright, D. M. (2009). Impact of deer invasion on New Zealand forests. PhD Thesis, University of Cambridge, Cambridge.
  • Wright, S., Zeballos, H., Dominguez, I., Gallardo, M., Moreno, M. & Ibanez, R. (2000). Poachers alter mammal abundance, seed dispersal, and seed predation in a Neotropical forest. Conservation Biology 14, 227239.
  • Zhang, C., Tian, H., Pan, S., Liu, M., Lockaby, G., Schilling, E. & Stanturf, J. (2008). Effects of forest regrowth and urbanization on ecosystem carbon storage in a rural–urban gradient in the southeastern United States. Ecosystems 11, 12111222.
  • Zhao, H., Zhao, X., Zhou, R., Zhang, T. & Drake, S. (2005). Desertification processes due to heavy grazing in sandy rangeland, Inner Mongolia. Journal of Arid Environments 62, 309319.