The main remnant of the head of the syrphid larva hag been retracted into the thorax and is called the cephalopharyngeal apparatus. It forms a special food collecting and sorting pump. The form, development, and function of this structure in Eristalis is described. The structure of the other remnants of the head is also considered.
The cephalopharyngeal apparatus of other Syrphidae is compared with the saprophagous type of Eristalis. A previously undescribed feature is a grinding mill in the cephalopharyngeal apparatus of Merodon and Eumerus. The way in which the various forms of the cephalopharyngeal apparatus of the Syrphidae have arisen from the primitive type is discussed.
The morphology of the head is compared with other families of Diptera and relationships of some parts are discussed. A new interpretation of some parts of the cephalopharyngeal apparatus is suggested, particularly the so-called ‘ intermediate ’ sclerite.
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