• Baker, P., Funk, S., Harris, S. & White, P. (2000). Flexible spatial organization of urban foxes, Vulpes vulpes, before and during an outbreak of sarcoptic mange. Anim. Behav. 59, 127146.
  • Barnard, P. (1998). Biological diversity in Namibia: a country study. 339pp. Namibian biodiversity task force. Windhoek.
  • Beyer, H. (2004). Hawth's Analysis Tools for ArcGIS. Available at (accessed 20 March 2006)
  • Burt, W. (1943). Territoriality and home range concepts as applied to mammals. J. Mammal. 24, 346352.
  • Dreyer, H.V.A. & Nel, J. (1990). Feeding site selection by black-backed jackals on the Namib Desert coast. J. Arid Ecol. 19, 217224.
  • Ferguson, J., Nel, J.A.J. & de Wet, M. (1983). Social organization and movement patterns of black-backed jackals Canis mesomelas in South Africa. J. Zool. (Lond.) 199, 487502.
  • Fretwell, S. & Lucas, H. (1970). On territorial behaviour and other factors influencing habitat distribution in birds. Acta Biotheor. 19, 1636.
  • Geffen, E., Gompper, M., Gittleman, J., Luh, H.-K., Dw, M. & Wayne, R. (1996). Size, life-history traits, and social organisation in the Canidae: a reevaluation. Am. Nat. 147, 140160.
  • Gowtage-Sequeira, S. (2005). The importance of jackals and domestic dogs for the transmission of generalist canid pathogens to sympatric carnivores in Namibia. PhD thesis, University of Edinburgh, Edinburgh
  • Harris, S., Cresswell, W.J., Forde, P.G., Trewhella, W.J., Woodland, T. & Wray, S. (1990). Home range analysis using radiotracking data – A review of problems and techniques particularly as applied to the study of mammals. Mammal. Rev. 20, 97123.
  • Hiscocks, K. & Perrin, M. (1987). Feeding observations and diet of black-backed jackals in an arid coastal environment. S. Afr. J. Wildl. Res. 17, 5558.
  • Hiscocks, K. & Perrin, M. (1988). Home range and movements of black-backed jackals at Cape Cross Seal Reserve, Namibia. S. Afr. J. Wildl. Res. 18, 97100.
  • Hofer, H. & East, M.L. (1993a). The commuting system of Serengeti spotted hyaenas: how a predator copes with migratory prey. I. Social organization. Anim. Behav. 46, 547557.
  • Hofer, H. & East, M.L. (1993b). The commuting system of Serengeti spotted hyaenas: how a predator copes with migratory prey. II. Intrusion pressure and commuters' space use. Anim. Behav. 46, 559574.
  • Höner, O., Wachter, B., East, M.L., Runyoro, V. & Hofer, H. (2005). The effect of prey abundance and foraging tactics on the population dynamics of a social, territorial carnivore, the spotted hyena. Oikos 108, 544554.
  • Jenner, N. (2008). Contributions to offspring care by parents and helpers in the black-backed jackal (Canis mesomelas). PhD thesis, Zoological Society of London's Institute of Zoology, London and University of Kent, Canterbury, UK
  • Jennions, M.D. & Macdonald, D. (1994). Cooperative breeding in mammals. Trends Ecol. Evol. 9, 8993.
  • Kruuk, H. & Macdonald, D.W. (1985). Group territories of carnivores: empires and enclaves. In Behavioural ecology: ecological consequences of adaptive behaviour: 521536. Sibly, R.M. & Smith, R.H. (Eds). Oxford: Blackwell Scientific Publications.
  • Loveridge, A.J. & Nel, J.A.J. (2004). Black backed jackal Canis mesomelas Schreber, 1775. In Status survey and conservation action plan – canids: foxes, wolves, jackals and dogs: 161166. Sillero-Zubiri, C., Hoffmann, M. & Macdonald, D.W. (Eds). Cambridge: IUCN/SSC Canid Specialist Group.
  • Macdonald, D. (1983). The ecology of carnivore social behaviour. Nature 301, 379384.
  • Maher, C. & Lott, D. (2000). A review of ecological determinants of territoriality within vertebrate species. Am. Midl. Nat. 143, 129.
  • Moehlman, P. (1979). Jackal helpers and pup survival. Nature 277, 382383.
  • Moehlman, P.D. (1983). Socioecology of silverbacked and golden jackals (Canis mesomelas and Canis aureus). In Advances in the study of mammalian behavior: 423451. Eisenberg, J.F. & Kleiman, D.G. (Eds). Pittsburg: American Society of Mammalogists.
  • Nel, J.A.J. & Loutit, R. (1986). The diet of the black-backed jackals, Canis mesomelas, on the Namib desert coast. Cimbebasia 8, 9196.
  • Rose, M. & Polis, G. (1998). The distribution and abundance of coyotes: the effects of allochthonous food subsidies from the sea. Ecology 79, 9981007.
  • Sillero-Zubiri, C. & Macdonald, D.W. (1998). Scent-marking and territorial behaviour of Ethiopian wolves Canis simensis. J. Zool. (Lond.) 245, 351361.
  • Wolff, J. & Peterson, J. (1998). An offspring-defence hypothesis for territoriality in female mammals. Ethol. Ecol. Evol. 10, 227239.