• (With Plates 3 and 4 and 2 figures in the text)


  • 1The sequence of events in the development of the inflorescence and spikelets of Anthoxanthum odoratum is described and illustrated.
  • 2The shoot-apex becomes longer and bears more leaf primordia during spring, becoming extremely elongated just prior to inflorescence initiation.
  • 3Primary panicle buds are borne in two rows in the axils of alternate small collar-like leaf primordia.
  • 4These buds bear rudimentary leaf primordia in whose axils secondary buds are initiated. Similar repeated branching leads to the development of the panicle type of inflorescence.
  • 5Spikelet differentiation begins at the tip of the inflorescence and later occurs at the ends of the individual panicle branches.
  • 6The two glumes and the four lemmas arise in an orderly alternate sequence, followed by the two stamen primordia.
  • 7The stamen primordia appear to be axial in nature and are not leaf primordia.
  • 8The two stamens are probably the axillary buds of the last two lemmas.
  • 9The carpel arises on the side of the spikelet apex in the plane of the lemmas and glumes, continuing their strictly alternate sequence.
  • 10In origin the carpel appears to be homologous with a single leaf.
  • 11The two stigmas arise one on each side of the carpel as the latter encloses the spikelet tip.
  • 12In the terminal spikelet the carpel appears to be the last leaf on the whole shoot and is in strict distichous continuity with (a) the lemmas and glumes of that spikelet, (b) the collar-like rudimentary leaf primordia subtending the primary spikelet-branches below, and (c) the vegetative leaves at the base.