Etyid crabs (Crustacea, Decapoda) from mid-Cretaceous Reefal strata of Navarra, northern Spain



Abstract:  All known etyid crabs (Crustacea, Decapoda) from the upper Albian – lower Cenomanian Aldoirar coralgal-dominated patch reef (Albeniz Unit, Eguino Formation), as exposed at the disused Koskobilo quarry in Navarra, northern Spain, are described, discussed and illustrated. A new species, Xanthosia koskobiloensis, the first member of the genus on record from southern Europe, is erected, and a variety that appears closely related to this, Xanthosia cf. X. koskobiloensis, is recognised. This new taxon could be a forerunner of the early Palaeocene (Danian) Xanthosia gracilis from Fakse (Sjælland, eastern Denmark), with which it shares a morphologically closely similar carapace. On the basis of a revised overview here of all species assigned to it, the genus Xanthosia may have evolved in an environment dominated by deposition of siliciclastics, rather than chalks. In addition, Etyxanthosia fossa has been collected at Koskobilo, and as the distribution of all known specimens demonstrates, E. fossa inhabited various environments. Another new species, Caloxanthus paraornatus, is closely similar as well to a species from Fakse, Caloxanthus ornatus, and constitutes the first record of the genus for southern Europe.