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Taxonomic studies of Pterostichus ishikawai Nemoto (Coleoptera: Carabidae)


Laboratory of Forest Zoology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, 1-1-1 Yayoi, Bunkyo-ku, Tokyo, 113-8657 Japan. Email:


A macrocephalic pterostichine carabid, Pterostichus ishikawai Nemoto, endemic to the Korean Peninsula, is taxonomically revised based mainly on the membranous parts of the genitalia (male endophallus and female vagina). Two species are separated from P. ishikawai (type locality: Mount Taebaegsan) and newly described: P. ishikawaioides (type locality: Mount Sobaeksan) and P. jiricola (type locality: Mount Jirisan). Since comparative male genital morphology does not support monophyly for these three species, the two novel species should be treated as distinct species, not subspecies of P. ishikawai.