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The genetic analysis of two abnormal characters, exlra sex combs and abnormal head shape in hybrids between Drosophila madeirensu and Drosophila subobscura. revealed a major effect of lite X chromosome in both cases. Autosomes also play a role in determining these abnormalities, (he E chromosome in case of extra sex combs, the E and O chromosomes in case of abnormal head shape. Autosomes do not cause Che Two abnormal character without interacting with the X chromosome.