Seeds of 14 species were collected, usually in each of 2 or 3 yr, and mixed with the top 7.5 cm of sterilised soil confined in cylinders sunk in the ground outdoors and cultivated three times yearly. Seedling emergence was recorded for at least 3 and usually 5 yr. Most seedlings of Ranunculus bulbosus and Rumex acetosa appeared during the autumn of sowing, while the main emergence of Geranium pratense. Ranunculus acris, Rumex conglomeratus and R. maritimus was in the following spring. Emergence of G. pyrenaicum and G. robertianum took place throughout much of the year after sowing. Seeds of all these species were relatively short-lived in cultivated soil, and few seedlings appeared after the second year.

Seed survival was greater in G. dissectum. G. molle, Rumex acetosella. Ranunculus sceleratus and especially R. repens, of which emergence in the fifth year after sowing amounted to 4% of the seeds sown. Emergence from the persistent seed bank was mainly from May to September for G. dissectum and G. molle and during autumn for Ranunculus flammula and R. sceleratus, while seedlings of Ranunculus repens and Rumex acetosella appeared sporadically over most of the year.