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Brazilian species of the genus Epidendrum, distributed over various habitats, were analysed for epidermal structure to test correlations between habitat form and variation in stomatal parameters. This study confirmed the tendency for stomatal index values to increase in the apical direction of the foliar surface in this genus. The generally held view that there are a smaller number of stomata on the adaxial surface in amphistomatic leaves was contradicted for E. vesicatum, whose pendent sympodia showed that the adaxial foliar surfaces assume the abaxial position in relation to light incidence.