• Alexandersen, S., U. Wernery, P. Nagy, T. Frederiksen, and P. Normann, 2008: Dromedaries (Camelus dromedarius) are of low susceptibility to inoculation with foot-and-mouth disease virus serotype O. J. Comp. Pathol. 139, 187193.
  • van Boheemen, S., M. de Graaf, C. Lauber, T.M. Bestebroer, V.S. Raj, A.M. Zaki, A.D. Osterhaus, B.L. Haagmans, A.E. Gorbalenya, E.J. Snijder, and R.A. Fouchier, 2012: Genomic characterization of a newly discovered coronavirus associated with acute respiratory distress syndrome in humans. MBio. 3, e0047312.
  • Butler, D., 2013: Progress stalled on coronavirus. Nature 501, 294295.
  • Frederiksen, T., J. Borch, J. Christensen, K. Tjornehoj, U. Wernery, and S. Alexandersen, 2006: Comparison of FMDV Type O, A and Asia 1 Antibody Levels After Vaccination of Cattle, Sheep, Dromedary Camels and Horses. Session of the Research Group of the Standing Technical Committee of EuFMD, Paphos, Cyprus, 17-20 October 2006 Appendix 36, pp. 243249. Available at: (accessed December 28, 2013).
  • Haagmans, B.L., S.H. Al Dhahiry, C.B. Reusken, V.S. Raj, M. Galiano, R. Myers, G.J. Godeke, M. Jonges, E. Farag, A. Diab, H. Ghobashy, F. Alhajri, M. Al-Thani, S.A. Al-Marri, H.E. Al Romaihi, A. Al Khal, A. Bermingham, A.D. Osterhaus, M.M. Alhajri, and M.P. Koopmans, 2013: Middle East respiratory syndrome coronavirus in dromedary camels: an outbreak investigation. Lancet Infect. Dis. 12, pii: S14733099.
  • Hawkes, N., 2013: Camels could be the source of MERS coronavirus, research finds. BMJ 347, f5052.
  • Hemida, M., R. Perera, P. Wang, M. Alhammadi, L. Siu, M. Li, L. Poon, L. Saif, A. Alnaeem, and M. Peiris, 2013: Middle East Respiratory Syndrome (MERS) coronavirus seroprevalence in domestic livestock in Saudi Arabia, 2010 to 2013. Euro. Surveill. 18, pii: 20659.
  • Kobinger, G.P., J.M. Figueredo, T. Rowe, Y. Zhi, G. Gao, J.C. Sanmiguel, P. Bell, N.A. Wivel, L.A. Zitzow, D.B. Flieder, R.J. Hogan, and J.M. Wilson, 2007: Adenovirus-based vaccine prevents pneumonia in ferrets challenged with the SARS coronavirus and stimulates robust immune responses in macaques. Vaccine 25, 52205231.
  • Kupferschmidt, K., 2013: Emerging diseases. Researchers scramble to understand camel connection to MERS. Science 341, 702.
  • Meyer, B., M.A. Müller, V.M. Corman, C.B.E.M. Reusken, D. Ritz, G.-J. Godeke, E. Lattwein, S. Kallies, A. Siemens, J. van Beek, J.F. Drexler, D. Muth, B.-J. Bosch, U. Wernery, M.P.G. Koopmans, R. Wernery, and C. Drosten, 2014: Antibodies against MERS coronavirus in dromedary camels, United Arab Emirates, 2003 and 2013. Emerg. Infect. Dis. [Internet]. Available at: (accessed January 2, 2014).
  • Perera, R.A., P. Wang, M.R. Gomaa, R. El-Shesheny, A. Kandeil, O. Bagato, L.Y. Siu, M.M. Shehata, A.S. Kayed, Y. Moatasim, M. Li, L.L. Poon, Y. Guan, R.J. Webby, M.A. Ali, J.S. Peiris, and G. Kayali, 2013: Seroepidemiology for MERS coronavirus using microneutralisation and pseudoparticle virus neutralisation assays reveal a high prevalence of antibody in dromedary camels in Egypt. Euro. Surveill. 18, pii: 20574.
  • Reusken, C., M. Ababneh, V. Raj, B. Meyer, A. Eljarah, S. Abutarbush, G. Godeke, T. Bestebroer, I. Zutt, M. Muller, B. Bosch, P. Rottier, A. Osterhaus, C. Drosten, B. Haagmans, and M. Koopmans, 2013a: Middle East Respiratory Syndrome coronavirus (MERS-CoV) serology in major livestock species in an affected region in Jordan, June to September 2013a. Euro. Surveill. 18, pii: 20662.
  • Reusken, C.B., B.L. Haagmans, M.A. Muller, C. Gutierrez, G.J. Godeke, B. Meyer, D. Muth, V.S. Raj, L.S. De Vries, V.M. Corman, J.F. Drexler, S.L. Smits, Y.E. El Tahir, R. De Sousa, J. van Beek, N. Nowotny, K. van Maanen, E. Hidalgo-Hermoso, B.J. Bosch, P. Rottier, A. Osterhaus, C. Gortazar-Schmidt, C. Drosten, and M.P. Koopmans, 2013b: Middle East respiratory syndrome coronavirus neutralising serum antibodies in dromedary camels: a comparative serological study. Lancet Infect. Dis. 13, 859866.
  • Wernery, U., P. Nagy, C.M. Amaral-Doel, Z. Zhang, and S. Alexandersen, 2006: Lack of susceptibility of the dromedary camel (Camelus dromedarius) to foot-and-mouth disease virus serotype O. Vet. Rec. 158, 201203.
  • WHO 2013: WHO Global Alert and Response (GAR): Middle East respiratory coronavirus – update 27 December 2013. (accessed December 27, 2013).
  • de Wit, E., and V.J. Munster, 2013: MERS-CoV: the intermediate host identified?. Lancet Infect Dis. 13, 827828.
  • Wong, G., J.S. Richardson, S. Pillet, A. Patel, X. Qiu, J. Alimonti, J. Hogan, Y. Zhang, A. Takada, H. Feldmann, and G.P. Kobinger, 2012: Immune parameters correlate with protection against ebola virus infection in rodents and nonhuman primates. Sci. Transl. Med. 4, 158ra146.
  • Zaki, A.M., S. van Boheemen, T.M. Bestebroer, A.D. Osterhaus, and R.A. Fouchier, 2012: Isolation of a novel coronavirus from a man with pneumonia in Saudi Arabia. N. Engl. J. Med. 367, 18141820.