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M. D. HOLDEN, J. GREGORY, V. WATKINS and L. RADFORD Operant-conditioning programme for White rhinoceros, Black rhinoceros and Indian or Greater one-horned Asian rhinoceros Ceratotherium simum, Diceros bi cornis and Rhinoceros unicornis at Whipsnade Wild Animal Park, Dunstable, UK International Zoo Yearbook 40

Version of Record online: 22 JUN 2006 | DOI: 10.1111/j.1748-1090.2006.00144.x

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