Parasite Immunology

Cover image for Vol. 36 Issue 12

Edited By: Eleanor M. Riley and Richard K. Grencis

Impact Factor: 1.849

ISI Journal Citation Reports © Ranking: 2013: 18/36 (Parasitology); 118/144 (Immunology)

Online ISSN: 1365-3024

Author Guidelines

If you have a query please contact the Managing Editor at

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  • Journal overview

Manuscript Submission

All papers must be submitted via ScholarOne Manuscripts.

Authors may wish to consult an appropriate member of the Editorial Board prior to submission of material.

On submission you will need: (1) the type of your paper (original paper, brief definitive report, Review) (2) the full title of your paper (limited to 120 characters), (3) the abstract (limited to 200 words), (4) up to six keywords, (5) the full names and affiliations of all authors, (6) details of preferred and/or non-preferred reviewers (optional), (7) a cover letter, (8) a manuscript keyword, (9) to answer questions regarding the Woods Hole Immuno-Parasitology Prize, human tissues and animal experimentation, (10) to complete and upload a Conflict of Interest form for each author, (11) to adhere to the ARRIVE guidelines, and (12) to upload the appropriate files and forms.

Submissions should be prepared using double-line spacing, a minimum of 1 inch margins, and 12 point Times New Roman font. The preferred file type formats for the text and tables of your manuscript are .doc, .docx, .rtf, .ppt, .xls; please do not save them as portable document files (PDF) files. Ideally, illustrations should be uploaded in native format of PICT if created on a Mac, or in native format or WMF if created in Windows. Files saved as PS, EPS, GIF and TIF may also be used. Avoid using tints if possible; if they are essential to the understanding of the figure, try to make them coarse. Detailed information on our digital illustration standards is available on-line at:

Each submission should include a title page which should contain: the paper title, full names and affiliations of all authors, and should indicate an author for use for correspondence (including an email address). For each author, disclose potential conflicts of interest, including all relevant financial interests (e.g. employment, significant share ownership, patent rights, consultancy, research funding) in any company or institution that might benefit from the publication (or state 'none'). Authors do not need to report the sums concerned. Please can you therefore add a small paragraph entitled "Disclosures" and then include any relevant details or "none" if there is nothing to disclose, if you have not already done so. For indexing purposes a short number of 'key words' should be included.

If you are not a native English-language speaker, we strongly recommend that you have your manuscript professionally edited before submission. A list of companies that will edit your manuscript for a fee can be found here Professional editing is not compulsory, but will mean that reviewers are better able to read and assess your manuscript. Use of one of these companies does not guarantee acceptance or preference for publication in this journal.

Manuscript Types and Style

Original Papers
The text should be preceded by a short abstract not exceeding 200 words. The abstract of a paper in Parasite Immunology is the face of the paper turned to the world. It is not a prècis of the paper, but is a clear, direct account of what was done, why it was done, and why the outcome matters. This is not the place for details or data, or for speculation. Avoid abbreviations, write sparingly, but simply and clearly as to a general audience, and give the highlights only. You can make some simple changes to your title and abstract to improve your article's ranking in search engines: guidelines on this can be found here. Please review your article's title and abstract in the light of these suggestions to improve your paper prior to submission and maximise its potential readership.

The abstract should be followed by these sections: Introduction, Materials and Methods, Results, Discussion, Acknowledgements, References, Legends to Tables and Figures, Tables and Figures. Pages should be numbered consecutively in arabic numerals. Please also ensure that the lines in your manuscript are numbered. Acknowledgements should include an indication of the source of funding for the work and the contribution of individual authors (see “Authorship” below).

References should be cited in the Vancouver style. In the text they should be numbered in superscript in the order in which they appear. The reference list should follow the numbering sequence used in the text and should include: the names and initials of all authors; the full title of the article; the source of reference using abbreviations for journal titles as shown in Index Medicus, the year, volume number and first and last pages. For references with more than six authors, the first three should be listed, followed by et al. For references cited from books, the title of the book should be followed by the names and initials of the editors, the edition, the place of publication, the publisher, the year of publication and the first and last pages.


1) Rogerson SJ, Novakovic S, Cooke BM, Brown GV. Plasmodium falciparum-infected erythrocytes adhere to the proteoglycan thrombomodulin in static and flow-base systems. Exp Parasitol 1997; 86 8-18.

2) Parks DR, Lanier LL, Herzenberg LA. In Handbook of Experimental Immunology, ed. Weir DM, Oxford: Blackwell Science Ltd; 1986; 29

Authors are responsible for the accuracy of their references.

Spelling should conform to The Concise Oxford Dictionary of Current English and units of measurement, symbols and abbreviations with those in Units, Symbols and Abbreviations (1977) published and supplied by the Royal Society of Medicine, 1 Wimpole Street, London W1M 8AE. This specifies the use of SI units.

Tables and figures should be referred to in the text together with an indication of their approximate position. Colour illustrations are encouraged; there is no charge to the author for the inclusion of colour illustrations.

Brief Definitive Reports
The journal publishes studies that are not required to be written up as conventional papers. They should not be structured like original papers but should contain brief accounts of background studies, methods used, results and discussion. References should be kept to a minimum and there should be no more than one table and one figure. Maximum length accepted is four pages of typescript. They will be given some priority in publication as the schedule permits.

Review Articles
We would be happy to receive Review articles with a limit of 5000 words that are structured in the same format as Original Articles.

Letters to the Editor
These are published if they comment usefully on material published in previous issues, and should not exceed 400 words.

Meeting Reports
Reports on meetings or parts of meetings concerned with parasite immunology will be published. Again, these reports will be commissioned, but the editors will be pleased to receive reports that might be used.


All authors must fulfil the following three criteria:
Have made a substantial contribution to research design, or the acquisition, analysis or interpretation of data;
Have drafted the paper or revised it critically;
Have approved the submitted and final versions.
In the Acknowledgments section of the paper all authors must indicate their specific contributions to the work described in the manuscript. Some examples include
X performed the research;
Y designed the research study;
Z contributed essential reagents or tools;
A analysed the data;
B wrote the paper.
An author may list more than one contribution, and more than one author may have contributed to the same element of the work. E.g. 'A performed the research, A and C analysed the data and wrote the paper, E contributed the knockout mice for the study and G designed the research study and wrote the paper'.

Supporting Information

Online Supporting Information can include additional explanatory notes, data sets, videos, lists, figures or tables that will not be published in the print edition of the journal and which are ancillary to, rather than central to, the article. Supporting Information must be approved by the Editors and should be supplied as a single PDF file headed by the title of the paper and the authors' names, addresses and contact information. Supporting Information will be published exactly as supplied and it is the author's responsibility to ensure that the material is logically laid out, adequately described, and in a format accessible to readers. Animations and other moving images or sound files in standard formats must be supplied as separate files. Figures and tables in Supporting Information should be referred to in the main text and labelled Fig. S1, Fig. S2 or Table S1, etc., in the order cited. Full guidelines and information on acceptable file formats may be found at

Display of Sequences

Prepare sequences as figures, not tables. This will ensure that proper alignment is preserved.

Submission of Sequences to GenBank 

Original DNA sequences reported in Parasite Immunology must also be submitted to GenBank. Instructions for submission can be found at the following address: An accession number should be supplied parenthetically at a relevant location in the text.

Human and Murine Genes

For human genes, use genetic notation and symbols approved by the HUGO Nomenclature Committee. Approved gene symbols should be obtained prior to submission from the HUGO Nomenclature Committee, For nomenclature guidelines, see White et al., 'Guidelines for Human Gene Nomenclature' [Genomics, 45, 468-471 (1997)]. The Gene Name Proposal form may be completed on the Nomenclature Web page: Use ISCN nomenclature for cytogenetics notation [Mitelman, F. (ed.) ISCN 1995: An International System for Human Cytogenetic Nomenclature, S. Karger, Basel]. Human gene names and loci should be written in  uppercase italics and Arabic numerals. Protein products are not italicized. For mouse strain and genetic nomenclature, refer to the International Committee on Standardized Genetic Nomenclature for Mice: New symbols and names for genes should be obtained prior to submission through the online symbol registry form at:

Microarray Databases

Parasite Immunology supports the efforts of the Microarray Gene Expression Data Society to standardize the presentation of microarray data, and we recommend that authors follow their guidelines and checklist ( In addition, the journal strongly recommends the supplemental microarray data be deposited in a public database such as Gene Expression Omnibus (or GEO, at or Array Express ( or submitted for peer-review with the initial submission of the manuscript.

Ethical Policy and Guidelines

Parasite Immunology encourages its contributors and reviewers to adopt the standards of the International Committee of Medical Journal Editors. The Editors reserve the right to reject a paper that does not meet these standards.

Parasite Immunology will not consider papers that have been accepted for publication or published elsewhere. Copies of existing manuscripts with potentially overlapping or duplicative material should be submitted together with the manuscript, so that the Editors can judge suitability for publication.

Please click here to read the Ethical Policies of Parasite Immunology.

For more detailed ethical guidelines please visit:


All papers to Parasite Immunology are checked for potential plagiarism using CrossCheck plagiarism detection software. By submitting your manuscript to Parasite Immunology you accept that your manuscript will be screened for plagiarism against previously published works.

Animal Experimentation

The Editors will not allow any papers to be published that describe experiments on living animals which may reasonably be presumed to have inflicted unnecessary pain or discomfort upon them. Experiments on living vertebrates or Octopus vulgaris should conform in principle to the legal requirements in the UK. Whenever appropriate, a statement should be included indicating that experiments were performed in accordance with local/national guidelines.

ARRIVE Guidelines

Authors are expected to comply with the ARRIVE guidelines for reporting research before submission of a manuscript. While the guidelines refer to animal experiments, most of the elements are common to all forms of research communication and adherence will strengthen the transparency and completeness of reporting.

Clinical Trials Registry

In accordance with the guidelines published by the International Committee of Medical Journal Editors (ICMJE), Parasite Immunology will require, as a condition of consideration for publication, that all clinical trials be registered in a public trials registry (for example, at Trials must register at or before the onset of patient enrollment. For more information, see and go to Section III.J.

Data Sharing

Parasite Immunology supports the efforts to encourage open sharing of publication-related data. Parasite Immunology adheres to the beliefs that authors should include in their publications the data, algorithms, or other information that is central or integral to the publication or make it freely and readily accessible; use public repositories for data whenever possible; and make patented material available under a license for research use. For more information, see the NAS website:

Distribution of Reagents

The Editors of Parasite Immunology have adopted the policy that any readily renewable resources mentioned in a journal article not already obtainable from commercial sources shall be made available to all qualified investigators in the field. The policy stems from the long-standing scientific principle that authenticity requires reproducibility. Publication in Parasite Immunology constitutes a de facto acceptance of this policy. Included are reagents that can be easily provided; specifically, nucleic acid sequences, cDNA and genomic clones, cell lines, and monoclonal antibody clones. Small amounts (sufficient for the replication of any in vitro work reported) of novel protein reagents are also considered easily transferable.

Although the Editors appreciate that many of the reagents mentioned in Parasite Immunology are proprietary or unique, neither condition is considered adequate grounds for deviation from this policy. Suitable material transfer agreements can be drawn up between the provider and requester, but if a reasonable request is turned down and submitted to the Editor-in-Chief, the corresponding author will be held accountable. The consequence for noncompliance is simple: the corresponding author will not publish in Parasite Immunology for the following three years.

Disclosures and Conflict Interest

Authors are required to disclose financial interests in any company or institution that might benefit from their publication. All authors must complete and sign the Conflict of Interests Form. The completed form must be returned to the editorial office. A competing interest exists when a primary interest (such as patients’ welfare or the validity of research) might be influenced by a secondary interest (such as financial gain or personal rivalry). It may arise for the authors of a Parasite Immunology article when they have a financial interest that may influence their interpretation of their results or those of others.

Financial interests are the easiest to define and they have the greatest potential to influence the objectivity, integrity or perceived value of a publication. They may include any or all, but are not limited to, the following:

• Personal financial interests: Stocks or shares in companies that may gain or lose financially through publication; consultant fees or fees from speakers bureaus other forms of remuneration from organisations that may gain or lose financially; patents or patent applications whose value may be affected by publication.
• Funding: Research support from organisations that might gain or lose financially through publication of the paper.
• Employment: Recent, present or anticipated employment of you or a family member by any organization that may gain or lose financially through publication of the paper. Any such competing interest that authors may have should be declared. The aim of the statement is not to eradicate competing interests, as they are almost inevitable. Papers will not be rejected because there is a competing interest, but a declaration on whether or not there are competing interests will be added to the paper.
• Patent rights
• Consultancy work. All authors must disclose competing interests, or state “none” via the Journal's ScholarOne Manuscripts website.

All sources of funding must be disclosed in the Acknowledgments section of the paper. List governmental, industrial, charitable, philanthropic and/or personal sources of funding used for the studies described in the manuscript. Attribution of these funding sources is preferred. Examples:

• This work was supported by a grant from the National Institutes of Health, USA (DKxxxx to AB).
• This work was supported by the Crohn's and Colitis Foundation of Canada (grant to AB and CD).
• This work was supported by a grant from Big Pharma Inc. (to AB) and equipment was donated by Small Pharma Inc. EF received a graduate studentship award from the University of xxxxx.
For papers where there are no competing interests, all authors must include the statement ‘Competing interests: the authors have no competing interests.’ We will also ask reviewers to provide a statement of competing interests.

Copyright and OnlineOpen

Authors of all accepted manuscripts are required to license copyright in their paper to Wiley-Blackwell.

OnlineOpen is available to authors of primary research articles who wish to make their article available to non-subscribers on publication, or whose funding agency requires grantees to archive the final version of their article. With OnlineOpen, the author, the author's funding agency, or the author's institution pays a fee to ensure that the article is made available to non-subscribers upon publication via Wiley Online Library, as well as deposited in the funding agency's preferred archive. Click here for the full list of terms and conditions.

Prior to acceptance there is no requirement to inform an Editorial Office that you intend to publish your paper OnlineOpen if you do not wish to. All OnlineOpen articles are treated in the same way as any other article. They go through the journal's standard peer-review process and will be accepted or rejected based on their own merit.

Papers are accepted on the understanding that they have not been and will not be published elsewhere.

If your paper is accepted, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper.

For authors signing the copyright transfer agreement

If the OnlineOpen option is not selected the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs below:

CTA Terms and Conditions

For authors choosing OnlineOpen

If the OnlineOpen option is selected the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):

Creative Commons Attribution License OAA

Creative Commons Attribution Non-Commercial License OAA

Creative Commons Attribution Non-Commercial -NoDerivs License OAA

To preview the terms and conditions of these open access agreements please visit the Copyright FAQs hosted on Wiley Author Services and visit

If you select the OnlineOpen option and your research is funded by The Wellcome Trust and members of the Research Councils UK (RCUK) you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with Wellcome Trust and Research Councils UK requirements. For more information on this policy and the Journal’s compliant self-archiving policy please visit:

Accepted Articles

'Accepted Articles' have been accepted for publication and undergone full peer review but have not been through the copyediting, typesetting, pagination and proofreading process. Accepted Articles are published online a few days after final acceptance, appear in PDF format only (without the accompanying full-text HTML) and are given a Digital Object Identifier (DOI), which allows them to be cited and tracked. The DOI remains unique to a given article in perpetuity. More information about DOIs can be found online at Given that Accepted Articles are not considered to be final, please note that changes will be made to an article after Accepted Article online publication, which may lead to differences between this version and the Version of Record. The Accepted Articles service has been designed to ensure the earliest possible circulation of research papers after acceptance.

Accepted articles will be indexed by PubMed; therefore the submitting author must carefully check the names and affiliations of all authors provided in the cover page of the manuscript, as it will not be possible to alter these once a paper is made available online in Accepted Article format. Subsequently the final copyedited and proofed articles will appear either as Early View articles in a matter of weeks or in an issue on Wiley Online Library; thelink to the article in PubMed will automatically be updated.

Citing Accepted Articles

To include the Digital Object Identifier (DOI) in a citation to an article, simply append it to the reference as in the following example:

Choi, B and Kropf, P. Evaluation of T cell responses in healing and nonhealing leishmaniasis reveals differences in T helper cell polarisation ex vivo and in vitro. Parasite Immunol; DOI: 10.1111/j.1365-3024.2008.01094.x

To link to an article from the author’s homepage, take the DOI (digital object identifier) and append it to " " as per the following example:

DOI 10.1111/pim.12023, becomes 10.1111/pim.12023


The corresponding author will receive an email alert containing a link to a web site. A working e-mail address must therefore be provided for the corresponding author. The proof can be downloaded as a PDF (portable document format) file from this site. Acrobat Reader will be required in order to read this file. This software can be downloaded (free of charge) from the following web site: Adobe Reader Download. This will enable the file to be opened, read on screen and printed out in order for any corrections to be added. Further instructions will be sent with the proof. Hard copy proofs will be posted if no e-mail address is available. Excessive changes made by the author in the proofs, excluding typesetting errors, will be charged separately.

Author Services

Author Services enables authors to track their article – once it has been accepted – through the production process to publication online and in print. Authors can check the status of their articles online and choose to receive automated e-mails at key stages of production. The author will receive an e-mail with a unique link that enables them to register and have their article automatically added to the system. Please ensure that a complete e-mail address is provided when submitting the manuscript. Visit Author Services for more details on online production tracking and for a wealth of resources including FAQs and tips on article preparation, submission and more.


Free access to the final PDF offprint or your article will be available via Author Services only. Please therefore sign up for Author Services if you would like to access your article PDF offprint and enjoy the many other benefits the service offers. Paper offprints may be ordered at prices quoted on the order form, which accompanies proofs, provided that the form is returned with the proofs. The cost is more if the order form arrives too late for the main print run. Offprints are normally dispatched within three weeks of publication of the issue in which the paper appears. Please contact the folowing if offprints do not arrive: C.O.S. Printers Pte Ltd., 9 Kian Teck Crescent, Singapore 628875; Fax: +65 6265 9074; Email: However, please note that offprints are sent by surface mail, so overseas orders may take up to six weeks to arrive. Electronic offprints are sent to the first author at his or her first email address on the title page of the paper, unless advised otherwise; therefore please ensure that the name, address and email of the receiving author are clearly indicated on the manuscript title page if he or she is not the first author of the paper.

Woods Hole Immuno-Parasitology Prize 

Parasite Immunology awards a prize for the best paper published each year by an early-stage investigator. An early-stage investigator is either a student (undergraduate, Masters or PhD level) or a research fellow with no more than three years of post-doctoral research experience, who is the first author of the paper. The prize winner will receive £500 to cover the cost of registering for the annual Woods Hole Immuno-Parasitology meeting and will also have the opportunity to present their paper at this meeting. On submission of your manuscript you will be asked to indicate whether the first author of your paper qualifies for consideration for the Prize.

Immunity, Inflammation and Disease

Parasite Immunology collaborates with Wiley’s open access journal Immunity, Inflammation and Disease to enable rapid publication of good quality research that we are unable to accept for publication in Parasite Immunology. Authors will be offered the option of having the paper, along with any related peer reviews, automatically transferred for consideration by the Editor of Immunity, Inflammation and Disease. Authors will not need to reformat or rewrite their manuscript at this stage, and publication decisions will be made a short time after the transfer takes place. The Editor of Immunity, Inflammation and Disease will accept submissions that report well-conducted research which reaches the standard acceptable for publication. Immunity, Inflammation and Disease is a Wiley Open Access journal and article publication fees apply. For more information please go to