Phycological Research

Cover image for Vol. 65 Issue 2

Edited By: Kazuhiro Kogame

Impact Factor: 1.42

ISI Journal Citation Reports © Ranking: 2015: 56/104 (Marine & Freshwater Biology)

Online ISSN: 1440-1835

Author Guidelines


The journal is published in an online-only format since 2015.

Print subscription and single issue sales are available from Wiley’s Print-on-Demand Partner. To order online click on ‘Subscribe/Renew’ on the journal’s homepage on the Wiley Online Library.


A member of the Japanese Society of Phycology will receive a print journal as part of the membership.


Thank you for your interest in Phycological Research. Please consult the following instructions for help in preparing your manuscript, and feel free to consult us with any questions. To ensure fast peer review and publication, manuscripts that do not adhere to the following instructions will be returned to the corresponding author for technical revision before undergoing peer review. We look forward to your submission.

If you prefer to download the Author Guidelines as PDF file, please click here.


Phycological Research is published by the Japanese Society of Phycology (JSP) and complements the Japanese Journal of Phycology. The Journal publishes international, basic or applied, peer-reviewed research dealing with all aspects of phycology including ecology, taxonomy and phylogeny, evolution, genetics, molecular biology, biochemistry, cell biology, morphology, physiology, and new techniques, to facilitate the international exchange of results.


The acceptance criteria for all papers are the quality and originality of the research and its significance to our readership. Except where otherwise stated, manuscripts are peer reviewed by at least two anonymous referees, an Associate Editor and the Editor-in-Chief. The Editorial Board reserves the right to refuse any material for publication and advises that authors should retain copies of submitted manuscripts and correspondence as these materials cannot be returned. Final acceptance or rejection rests with the Editor-in-Chief. Where contributions are judged as acceptable for publication on the basis of scientific content, the Editor-in-Chief or the Publisher reserves the right to modify text to eliminate ambiguity and repetition and improve communication between author and reader.

Committee on Publication Ethics. The journal is a member of, and subscribes to the principles of the Committee on Publication Ethics (COPE).

Plagiarism Detection. The journal employs a plagiarism detection system. By submitting your manuscript to this journal you accept that your manuscript may be screened for plagiarism against previously published works.


Accepted papers will be passed to Wiley’s production team for publication. The author identified as the formal corresponding author for the paper will receive an email prompting them to login into Wiley’s Author Services, where via the Wiley Author Licensing Service (WALS) they will be asked to complete an electronic license agreement on behalf of all authors on the paper.

Authors may choose to publish under the terms of the journal’s standard copyright transfer agreement (CTA), or under open access terms made available via Wiley OnlineOpen.

Standard Copyright Transfer Agreement: FAQs about the terms and conditions of the standard CTA in place for the journal, including standard terms regarding archiving of the accepted version of the paper, are available at: Copyright Terms and Conditions FAQs.

OnlineOpen – Wiley’s Open Access Option: OnlineOpen is available to authors of articles who wish to make their article freely available to all on Wiley Online Library under a Creative Commons license. In addition, authors of OnlineOpen articles are permitted to post the final, published PDF of their article on their personal website, and in an institutional repository or other free public server immediately after publication. With OnlineOpen, the author, the author's funding agency, or the author's institution pays a fee to ensure that the article is made open access—known as “gold road” open access.

OnlineOpen licenses. Authors choosing OnlineOpen retain copyright in their article and have a choice of publishing under the following Creative Commons License terms: Creative Commons Attribution License (CC BY); Creative Commons Attribution Non-Commercial License (CC BY NC); Creative Commons Attribution Non-Commercial-NoDerivs License (CC BY NC ND). To preview the terms and conditions of these open access agreements please visit the Copyright Terms and Conditions FAQs.

If you select the OnlineOpen option and your research is funded by certain funders [e.g. The Wellcome Trust and members of the Research Councils UK (RCUK) or the Austrian Science Fund (FWF)] you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with your Funder requirements.

Funder Open Access and Self-Archiving Compliance: Please click here for more information on Wiley’s compliance with specific Funder Open Access and Self Archiving Policies, and click here for more detailed information especially about Self-Archiving definitions and policies.


Optimising Your Article for Search Engines
Many students and researchers looking for information online will use search engines such as Google, Yahoo or similar. By optimising your article for search engines, you will increase the chance of someone finding it. This in turn will make it more likely to be viewed and/or cited in another work. We have compiled these guidelines to enable you to maximise the web-friendliness of the most public part of your article.

Pre-acceptance English-language editing
Non-English speakers should have their manuscripts checked by a competent English speaker before submission. Manuscripts in which the English is difficult to understand may be returned to the authors for revision before scientific review. A list of independent providers of editing services can be found at All services are paid for and arranged by the author, and use of one of these services does not guarantee acceptance or preference for publication.

Manuscripts Categories and Word Limits
Contributions may take the form of Review Articles, Original Research Articles, Research Notes, and Book Reviews. Original Research Articles are limited to 8 printed pages. Research Notes should not normally exceed 4 printed pages. Approximately three typed pages will reduce to about one printed page. Figures and Tables are counted towards the page limit. Authors wishing to contribute Review Articles should contact the Editor or Associate Editors before submission.

Page charges for excess pages: For papers exceeding the above page limits, a page charge of ¥10,000 or (¥5,000 if the first or corresponding author is a JSP member) per additional printed page is levied to the author by JSP.

Formatting and Style

  • Document formatting. The main document should be saved in an editable format (e.g., MS-Word, RTF). Lines should be double-spaced, including references, tables and figure legends. Times New Roman font in 12-point type is preferred. Do not right-justify the text or divide words at the ends of lines. All pages and lines should be numbered serially to facilitate the review/editing processes.
  • Measurements. All measurements must be given in SI units as outlined in the latest edition of Units, Symbols and Abbreviations.
  • Abbreviations should be used sparingly and only where they ease the reader's task by reducing repetition of long, technical terms. Initially use the word in full, followed by the abbreviation in parentheses. Thereafter use the abbreviation.
  • Scientific names. The complete scientific names (genus, species, and authority), and cultivar or strain where appropriate, should be given for every organism when first mentioned. Common names of organisms, if used, must be accompanied by the correct scientific name on first mention. New botanical names proposed in the manuscript must also be mentioned in the Summary.
  • Footnotes to the text are not allowed and any such material should be incorporated into the text as parenthetical matter.

Parts of the manuscript

(i) Title page

The title page should contain (i) the title of the paper (including the class/division designation when a generic or specific name is used, but not containing authorities for scientific names), (ii) the full names of the authors, (iii) the addresses of the institutions at which the work was carried out, as well as the present address of any author if different to that where the work was carried out, (iv) the full postal and email address, plus fax and telephone numbers of the author to whom correspondence regarding the manuscript, proofs, and requests for offprints should be sent. The title should be short, informative and contain the major key words. A short running title (less than 40 characters, including spaces) should also be provided.

(ii) Summary and key words

Articles must have a Summary that states in 300 words or less, the purpose, basic procedures, main findings and principal conclusions of the study. The names of organisms used should be given, and new taxa that are described should be mentioned. Authorities for taxon names should not be given in the abstract unless the primary purpose of the paper is to clarify nomenclature and the authorship of taxa. The Summary should not contain abbreviations or references. Three to ten key words listed in alphabetical order should be supplied below the abstract for the purposes of indexing; note that words included in the title should not be included in key words.

(iii) Text

Introduction: This section should include sufficient background information to place the work in context. The aims of the manuscript should be clearly stated. The introduction should not contain either findings or conclusions.

Materials and methods: This should be concise but provide sufficient detail to allow the work to be repeated by others. The authors should include the sources of equipment, reagents and software (model number, company name, city, country when first referenced). Locations of sampling or study sites should be given using latitude and longitude coordinates. Authors should deposit voucher materials, strains, molecular sequences, and phylogenetic datasets in appropriate accession services (e.g., herbaria, culture collections, GenBank, TreeBASE, etc.).

Results: Results should be presented in a logical sequence in the text, tables and figures; repetitive presentation of the same data in different forms should be avoided. The Results should not contain material that would be appropriate for the Discussion.

Discussion: This should consider the results in relation to any hypotheses advanced in the Introduction and place the study in the context of other work. Only in exceptional cases should the Results and Discussion sections be combined.

(iv) Acknowledgments

The source of financial grants and other funding should be acknowledged, including a frank declaration of the authors' industrial links and affiliations. Do not acknowledge anonymous referees.

(v) References

In the Text: In the text, give the author's name followed by the year in parentheses: Sago (2000). If there are two authors, use 'and': Baskin and Baskin (1998); but if cited within parentheses, use '&': (Baskin & Baskin 1998). When reference is made to a work by three or more authors, the first name followed by et al. should be used: Powles et al. (1998). If several references by the same author(s) and from the same year are cited, a, b, c etc. should be put after the year of publication. Within parentheses, groups of references should be listed in chronological order. Personal communication, unpublished data and publications from informal meetings are not to be listed in the reference list but should be listed in full in the text (e.g. Smith A, 2000, unpublished data).

In the List of References: In a list at the end of the paper, references should be arranged alphabetically by author and then date. Cite the names of all authors when there are seven or fewer, and if there are more than seven, cite the first three plus et al. References should be listed in the following form.


Sato, M., Sakayama, H., Sato, M., Ito, M. and Sekimoto, H. 2014. Characterization of sexual reproductive processes in Chara braunii (Charales, Charophyceae). Phycol. Res. 62: 214–21.
Komazawa, I., Sakanishi, Y. and Tanaka, J. 2014. Temperature requirements for growth and maturation of the warm temperate kelp Eckloniopsis radicosa (Laminariales, Phaeophyta). Phycol. Res. Published online: 10 Oct. 2014; DOI: 10.1111/pre.12068


South, G. R. and Whittick, A. 1987. An Introduction to Phycology. Blackwell Science, Oxford.

Chapter in a book

Wynne, M. J. 1981. Phaeophyta: Morphology and classification. In Lobban, C. S. and Wynne, M. J. (Eds) The Biology of Seaweeds. Blackwell Science, Oxford, pp. 52–85.


Guiry, M. D. and Guiry, G. M. 2014. AlgaeBase. World-wide electronic publication, National University of Ireland, Galway. [cited on 21 November 2014]. Available from:

(vi) Tables

Tables should be self-contained and complement, but not duplicate, information contained in the text. Number tables consecutively in the text with Arabic numerals. Each table should be provided on a separate page with the table legend provided above the table. Legends should be concise but comprehensive - the table, legend, and footnotes must be understandable without reference to the text. Vertical lines should not be used to separate columns. Column headings should be brief, with units of measurement in parentheses; all abbreviations must be defined in the footnotes. The footnote symbols †, ‡, §, ¶, should be used in the order given, and *, **, *** should be reserved for indicating the significance of P-values. Statistical estimates such as SD or SE should be identified in the headings.

(vii) Figure legends

Provide all of the figure legends, separate from the figures, in this section. Legends should be concise but comprehensive - the figure and its legend must be understandable without reference to the text. Include definitions of any symbols used and define/explain all abbreviations and units of measurement. Provide a letter stating authorization if figures have been reproduced from another source. Labels in figures and figure legends should be set in alphabetical lowercase (e.g., a, b, c).

(viii) Figures

Figures should be cited in consecutive order in the text. Do not embed figures in the document - they must be supplied in separate files. Figures should be sized to fit within a one column (80 mm), span the page width (168 mm), or be of intermediate width (110 mm). Each figure, in its final size, must be a separate TIFF, JPEG, or PNG file with a resolution of 300 dpi or higher. If your original image is less than 300 dpi, the figure must be re-created because you cannot increase the resolution by simply resaving it at 300 dpi. More details on submitting artwork can be found on the Web site at

Labels in figures: Labels in figures should be set in alphabetical lowercase (e.g., a, b, c).

Drawings, graphs: The labels (8 point or larger) and the outer line on bars or boxes (at least 1-point font) should be adequately sized in the final size. Axis labels should be of uniform font and size and should not be bolded. Use an initial capital letter for the first work of each label (e.g., Wavelength, not wavelength). If units are included in the label, use the SI style (e.g., µmol photons m-2 s-1). All species names should be italicized within the figure.

Photographs: Individual photographs used to generate a plate should be of equal contrast. Each photograph must contain a scale measure, and be separated from other photographs in the plate by thin white lines (ca. 1mm). Scale length may be indicated directly above the scale or in the legend.

Color figures: [Non-JSP members] The journal is now online only publication so figures will be published in color online free of charges. [JSP Members] You will receive a print copy of the journal as part of the membership. If you wish figures of your article printed in color, ¥70,000 for the first three color figure plates and ¥35,000 for each extra color figure plate thereafter will be charged to the author. If authors reproduce figures in color for the online version but in black and white for the print version (no color charge), they should liaise with the Publisher to ensure that the appropriate documentation is completed and also prepare figures to be detailed enough to convey the necessary information even after they are converted into black and white.

(ix) Supporting information

Supporting Information can be a useful way for an author to include important but ancillary information with the online version of an article. Supporting Information is not essential to the article, but provides greater depth and background, and may include additional tables, data sets, figures, movie files, audio clips, 3D structures, and other related nonessential multimedia files. Supporting Information should be cited within the article text, and a descriptive legend should be included. It is published as supplied by the author, and a proof is not made available prior to publication; for these reasons, authors should provide any Supporting Information in the desired final format. Please note that the provision of supplementary material is not encouraged as a general rule. It will be assessed critically by reviewers and editors and will only be accepted if it is essential.

For further information on recommended file types and requirements for submission, please visit:

Authors should include a ‘Supporting Information’ section immediately after their References section, which should be in the following form (*only short titles to appendices should be given in this section; full titles can be given with the Supporting Information itself):

Supporting Information

Additional Supporting Information may be found in the online version of this article at the publisher's web-site:

Appendix S1: Short title of supplementary appendix S1*

Appendix S2: Short title of supplementary appendix S2*

Research Notes

Research Notes should contain a summary and key words, however the remainder of the text does not need to follow the format described above. References, tables, and figures must follow the format described above.


Wiley Author Services: After acceptance

Author Services enables authors to track their article through the production process to publication online and in print. Authors can check the status of their articles online and choose to receive automated e-mails at key stages of production. Visit Visit Author Services ( for more details on online production tracking.


Proofs will be sent via email as a PDF file and should be returned within 3 days of receipt. Alterations to the text and figures (other than the essential correction of errors) are unacceptable at proof stage and authors may be charged for excessive alterations. Authors should therefore supply an email address to which proofs can be emailed.


A minimum of 50 offprints will be provided upon request, at the author's expense. An Offprint Order Form outlining the cost of offprints will be sent to the corresponding author with the page proofs. Offprints will be provided only if a completed Offprint Order Form is returned to the publisher by the specified date.


If you fail to order the offprints and would like to order reprints of your article after publication of an issue, such orders can be made online for a fee. Please click on the following link and fill in the necessary details and ensure that you type information in all of the required fields: If you have queries about offprints, please email:

Early View Service

Phycological Research is covered by our Early View service. Early View articles are complete full-text articles published online in advance of their publication in a printed issue. Articles are therefore available as soon as they are ready, rather than having to wait for the next scheduled print issue. Because they are in final form, no changes can be made after online publication. The nature of Early View articles means that they do not yet have volume, issue or page numbers, so Early View articles cannot be cited in the traditional way. They are therefore given a Digital Object Identifier (DOI), which allows the article to be cited and tracked before it is allocated to an issue. After print publication, the DOI remains valid and can continue to be used to cite and access the article. More information about DOIs can be found at


Professor Kazuhiro Kogame
Department of Biological Sciences
Faculty of Science, Hokkaido University
Sapporo, 060-0810 Japan

Author Guidelines updated January 2017