Phycological Research

Cover image for Vol. 62 Issue 3

Edited By: Mitsunobu Kamiya

Impact Factor: 0.955

ISI Journal Citation Reports © Ranking: 2013: 72/102 (Marine & Freshwater Biology)

Online ISSN: 1440-1835

Author Guidelines

Thank you for your interest in Phycological Research. Please consult the following instructions for help in preparing your manuscript, and feel free to consult us with any questions. To ensure fast peer review and publication, manuscripts that do not adhere to the following instructions will be returned to the corresponding author for technical revision before undergoing peer review. We look forward to your submission.

If you prefer to download the Author Guidelines as PDF file, please click here.

Aims and Scope

Phycological Research is published by the Japanese Society of Phycology and complements the Japanese Journal of Phycology. The Journal publishes international, basic or applied, peer-reviewed research dealing with all aspects of phycology including ecology, taxonomy and phylogeny, evolution, genetics, molecular biology, biochemistry, cell biology, morphology, physiology, and new techniques, to facilitate the international exchange of results.

Contributions may take the form of Review Articles, Original Research Articles, Research Notes, and Book Reviews. Original Research Articles are limited to eight (8) printed pages. Research Notes should not normally exceed four (4) printed pages. Authors wishing to contribute Review Articles should contact the Editor or Associate Editors before submission.


The acceptance criteria for all papers are the quality and originality of the research and its significance to our readership. Except where otherwise stated, manuscripts are peer reviewed by at least two anonymous referees, an Associate Editor and the Editor-in-Chief. The Editorial Board reserves the right to refuse any material for publication and advises that authors should retain copies of submitted manuscripts and correspondence as these materials cannot be returned. Final acceptance or rejection rests with the Editor-in-Chief.

Manuscripts should be written so that they are intelligible to the professional reader who is not a specialist in the particular field. Where contributions are judged as acceptable for publication on the basis of scientific content, the Editor-in-Chief or the Publisher reserves the right to modify text to eliminate ambiguity and repetition and improve communication between author and reader. If extensive alterations are required, the manuscript will be returned to the author for revision.

Plagiarism Detection. Phycological Research employs a plagiarism detection system. By submitting your manuscript to this journal you accept that your manuscript may be screened for plagiarism against previously published works.

Committee on Publication Ethics. Phycological Research is a member of, and subscribes to the principles of the Committee on Publication Ethics (COPE).


Author Services

Prior to submission, we encourage you to browse the ‘Author Resources’ section of the Wiley Author Services website:

This site includes useful information covering such topics as copyright matters, ethics, electronic artwork guidelines, and much more.

English Language Editing Service

Authors for whom English is a second language may choose to have their manuscript professionally edited before submission to improve the English. Visit to learn about the options. All services are paid for and arranged by the author.  Please note using the Wiley English Language Editing Service does not guarantee that your paper will be accepted by this journal, and all services are paid for and arranged by the author.

Optimizing Your Article for Search Engines

Many students and researchers looking for information online will use search engines such as Google, Yahoo or similar. By optimizing your article for search engines, you will increase the chance of someone finding it. This in turn will make it more likely to be viewed and/or cited in another work. We have compiled these guidelines to enable you to maximize the web-friendliness of the most public part of your article.


Cover letter

A cover letter should be included with the manuscript and the following information should be provided:

         A declaration that the paper is being submitted for consideration for publication in Phycological Research and that the content has not been published or submitted for publication elsewhere. Authors should declare any financial support or relationships that may pose a conflict of interest.

         All authors' agreement: acknowledgement that all authors have contributed significantly, and that all authors are in agreement with the content of the manuscript.

         Contact details where the corresponding author can be contacted, including mailing address, telephone and fax numbers, and email address.

         If tables or figures have been reproduced from another source, a letter from the copyright holder (usually the Publisher), stating authorization to reproduce the material, must be included with the cover letter.


If your paper is accepted, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper.

For authors signing the copyright transfer agreement

If the OnlineOpen option is not selected the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs below:

CTA Terms and Conditions right.asp

For authors choosing OnlineOpen

If the OnlineOpen option is selected the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):

Creative Commons Attribution License OAA

Creative Commons Attribution Non-Commercial License OAA

Creative Commons Attribution Non-Commercial -NoDerivs License OAA

To preview the terms and conditions of these open access agreements please visit the Copyright FAQs hosted on Wiley Author Services right.asp and visit

If you select the OnlineOpen option and your research is funded by The Wellcome Trust and members of the Research Councils UK (RCUK) you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with Wellcome Trust and Research Councils UK requirements. For more information on this policy and the Journal’s compliant self-archiving policy please visit:


All manuscripts should be printed and the lines should be double-spaced, including references, tables and figure legends. Times New Roman font in 12-point type is preferred. The top, bottom and side margins should be 30 mm in A4 paper size. Do not right-justify the text or divide words at the ends of lines. All pages and lines should be numbered serially to facilitate the review/editing processes.
• Do not use the carriage return (enter) at the end of lines within a paragraph.
• Turn the hyphenation option off.
• Specify any special characters used to represent non- keyboard characters.
• Take care not to use l (ell) for 1 (one), O (capital o) for 0 (zero) or ß (German esszett) for β (Greek beta).


The Journal uses American English spelling and authors should therefore follow the latest edition of Merriam-Webster's Collegiate Dictionary. All measurements must be given in SI units as outlined in the latest edition of Units, Symbols and Abbreviations: A Guide for Medical and Scientific Editors and Authors (Royal Society of Medicine Press, London).

Abbreviations should be used sparingly and only where they ease the reader's task by reducing repetition of long, technical terms. Initially use the word in full, followed by the abbreviation in parentheses. Thereafter use the abbreviation.

The complete scientific names (genus, species, and authority), and cultivar or strain where appropriate, should be given for every organism when first mentioned. The generic name may be abbreviated to an initial in subsequent references except where intervening references to other genera would cause confusion. Common names of organisms, if used, must be accompanied by the correct scientific name on first mention. New botanical names proposed in the manuscript must also be mentioned in the Summary.

Parts of the manuscript

Manuscript should consist of the “Main Document” including (i) Title page, (ii) Summary and key words, (iii) Text, (iv) Acknowledgment, (v) References, (vi) Tables, and (vii) Figure legends, and “Figures”.  Please note that we do not accept files saved in PDF, MS-PowerPoint or MS-Excel format, for the main document and figures.

(1) Main Document

A single file containing a (i) title page, (ii) summary and key words, (iii) text, (iv) acknowledgments, (v) references, (vi) tables (each table complete with title and footnotes), and (vii) figure legends should be saved in an editable format (e.g. MS-Word, RTF). Footnotes to the text are not allowed and any such material should be incorporated into the text as parenthetical matter.

(2) Figures

Separate image files for each figure (or composite figure) saved in an ordinary format (TIFF, JPEG, GIF, BMP, PNG...etc.) with a resolution high enough to allow adequate peer reviews should be uploaded. Please note that 300 dpi TIFF for halftone images and EPS or 800 dpi TIFF for line art or combination figures will be required after acceptance for publication.

(i) Title page

The title page should contain (a) the title of the paper (including the class/division designation when a generic or specific name is used, but not containing authorities for scientific names), (b) the full names of the authors, (c) the addresses of the institutions at which the work was carried out, as well as the present address of any author if different to that where the work was carried out, (d) the full postal and email address, plus fax and telephone numbers of the author to whom correspondence regarding the manuscript, proofs, and requests for offprints should be sent.

The title should be short, informative and contain the major key words. A short running title (less than 40 characters, including spaces) should also be provided.


(ii) Summary and key words

Articles must have a Summary that states in 300 words or fewer, the purpose, basic procedures, main findings and principal conclusions of the study. The names of organisms used should be given, and new taxa that are described should be mentioned. Authorities for taxon names should not be given in the abstract unless the primary purpose of the paper is to clarify nomenclature and the authorship of taxa. New botanical names proposed in the manuscript must also be mentioned in the Summary. The Summary should not contain abbreviations or references. Three to ten key words listed in alphabetical order should be supplied below the abstract for the purposes of indexing; note that words included in the title need not be included in key words.


(iii) Text

Authors should use the following subheadings to divide the sections of their manuscript.

         Introduction: This section should include sufficient background information to place the work in context. The aims of the manuscript should be clearly stated. The introduction should not contain either findings or conclusions.

         Materials and methods: This should be concise but provide sufficient detail to allow the work to be repeated by others. The authors should include the sources of equipment, reagents and software (model number, company name, city, country when first referenced). Locations of sampling or study sites should be given using latitude and longitude coordinates. Authors should deposit voucher materials, strains, molecular sequences, and phylogenetic datasets in appropriate accession services (e.g., herbaria, culture collections, GenBank, TreeBASE, etc.).

         Results: Results should be presented in a logical sequence in the text, tables and figures; repetitive presentation of the same data in different forms should be avoided. The Results should not contain material that would be appropriate for the Discussion.

         Discussion: This should consider the results in relation to any hypotheses advanced in the Introduction and place the study in the context of other work. Only in exceptional cases should the Results and Discussion sections be combined.


(iv) Acknowledgments

The source of financial grants and other funding should be acknowledged, including a frank declaration of the authors' industrial links and affiliations. The contribution of colleagues or institutions should also be acknowledged. Acknowledgement of anonymous referees is not permitted.


(v) References

The Harvard (author, date) system of referencing is used.

         In the Text:

In the text, give the author's name followed by the year in parentheses: Sago (2000). If there are two authors, use 'and': Baskin and Baskin (1998); but if cited within parentheses, use '&': (Baskin & Baskin 1998). When reference is made to a work by three or more authors, the first name followed by et al. should be used: Powles et al. (1998). If several references by the same author(s) and from the same year are cited, a, b, c etc. should be put after the year of publication. Within parentheses, groups of references should be listed in chronological order.

Personal communication, unpublished data and publications from informal meetings are not to be listed in the reference list but should be listed in full in the text (e.g. Smith A, 2000, unpublished data).

Please see the latest issues to clarify the writing convention in the text.


         In the List of References:

In a list at the end of the paper, references should be arranged alphabetically by author and then date. Cite the names of all authors when there are seven or fewer, and if there are more than seven, cite the first three plus et al. References should be listed in the following form.


o    Journals

Suzuki, M., Hashimoto, T., Nakayama, T. and Yoshizaki, M. 2010. Morphology and molecular relationships of Leptofauchea rhodymenioides (Rhodymeniales, Rhodophyta), a new record for Japan. Phycol. Res. 58: 11631.


o    Standard Journal article using DOI articles published online in advance without volume, issue, or page number (More information about DOIs:

Yu, J., Li, Z. and Brand J. J. 2009. Characterization of a green alga isolated from infected human external tissue Phycol.Res. Published online: December 7 2009; DOI: 10.1111/j.1440 1835.2009.00544.x


o    Books

South, G. R. and Whittick, A. 1987. An Introduction to Phycology. Blackwell Science, Oxford.


o    Chapter in a book

Wynne, M. J. 1981. Phaeophyta: Morphology and classification. In Lobban, C. S. and Wynne, M. J. (Eds) The Biology of Seaweeds. Blackwell Science, Oxford, pp. 5285.


o    Website

Ocean Primary Productivity Team (home page on the internet). 2000. Ocean primary productivity study. Institute of Marine Coastal Sciences, Rutgers University, New Brunswick [updated: 5 October 2000; cited 21 January 2002]. Available from:


(vi) Tables

Tables should be self- contained and complement, but not duplicate, information contained in the text. Number tables consecutively in the text with Arabic numerals. Each table should be provided on a separate page with the table legend provided above the table. Legends should be concise but comprehensive - the table, legend, and footnotes must be understandable without reference to the text. Vertical lines should not be used to separate columns. Column headings should be brief, with units of measurement in parentheses; all abbreviations must be defined in the footnotes. The footnote symbols †, ‡, §, ¶, should be used in the order given, and *, **, *** should be reserved for indicating the significance of P-values. Statistical estimates such as SD or SEM should be identified in the headings.
• Use a tab, not spaces, to separate data in tables.
• If you use a table editor function, ensure that each datum is contained within a unique cell; i.e. do not use carriage returns within cells.


(Vii) Figure legends

Provide all of the figure legends, separate from the figures, in this section. Legends should be concise but comprehensive – the figure and its legend must be understandable without reference to the text. Include definitions of any symbols used and define/explain all abbreviations and units of measurement. Provide a letter stating authorization if figures have been reproduced from another source.


(viii) Figures

All illustrations (line drawings and photographs) are classified as figures. Figures should be cited in consecutive order in the text. Do not embed figures in the document – they must be supplied in separate files. Line figures should be supplied as sharp, black and white graphs or diagrams, drawn professionally or with a computer graphics package. Lettering must be included and should be sized to be no larger than the journal text. Figures should be sized to fit within a one column (80 mm), span the page width (168 mm), or be of intermediate width (110 mm).


Regarding the format for digital figure files, 300 dpi TIFF for halftone images and EPS or 800 dpi TIFF for line art and combination of halftone and line art figures are recommended. Other common image file formats (e.g. JPEG, PNG) are also allowed, however it is the authors' responsibility to provide figures of appropriate quality. In case that the provided files have poor resolution or problematic, authors may be asked to resubmit the figures after acceptance.

Wiley Author Services also gives detailed information on the preparation and submission of articles at, and figures at

• Individual photographs forming a plate should be of equal contrast to facilitate printing.

• Magnifications should be indicated using a scale bar on the illustration.


Color figures Charges

A charge of ¥70,000 for the first three color figure plate(s) and ¥35,000 for each extra color figure plate thereafter will be charged to the author.


Color on Web Services

In the event that an author is not able to cover the costs of reproducing color figures in color in the printed version of the journal, Phycological Research offers authors the opportunity to reproduce color figures in color for free in the online version of the article (but they will still appear in black and white in the print version). If an author wishes to take advantage of this free color-on-the-web service, they should liaise with the Publisher to ensure that the appropriate documentation is completed. Please note that, in such cases, it is the authors' responsibility to prepare figures to be detailed enough to convey the necessary information even after they are converted into black and white.


Supporting Information

Supporting information is not essential to the article but provides greater depth and background and may include tables, figures, videos, datasets, etc. This material can be submitted with your manuscript, and will appear online, without editing or typesetting. Such supporting information should be referred to in the text as, for example, “see Appendix S1 in Supporting Information.” Guidelines on how to prepare this material and which formats and files sizes are acceptable can be found at:

Please note that the provision of supporting information is not encouraged as a general rule. It will be assessed critically by reviewers and editors and will only be accepted if it is essential. 

The author must advise Wiley-Blackwell if the URL of the web site where the Supporting Information is located changes. The content of the Supporting Information must not be altered after the paper has been accepted for publication.


Authors should include a ‘Supporting Information’ section immediately after their References section, which should be in the following form: 


Supporting Information

Additional Supporting Information may be found in the online version of this article:


Appendix S1 Short title of supplementary appendix S1*

Appendix S2 Short title of supplementary appendix S2*


Please note: Wiley is not responsible for the content or functionality of any supplementary materials supplied by the authors. Any queries (other than missing material) should be directed to the corresponding author for the article.


*Only short titles to appendices should be given in this section; full titles can be given with the Supporting Information itself.



Research Notes

Research Notes should contain a summary and keywords, however the remainder of the text does not need to follow the format described above. References, tables, and figures must follow the format describe above.



Manuscripts should be submitted electronically via ScholarOne Manuscripts at The use of an online submission and peer review site enables immediate distribution of manuscripts and consequentially speeds up the review process. It also allows authors to track the status of their own manuscripts. Complete instructions for submitting a paper is available at the ScholarOne Manuscripts site and below. Further assistance can be obtained from:


Getting Started

Launch your web browser (supported browsers include Internet Explorer 6 or higher, Netscape 7.0, 7.1, or 7.2, Safari 1.2.4, or Firefox 1.0.4) and go to the journal's online submission site:<xxx>. Log-in or click the 'Create Account' option if you are a first-time user.

Creating a New Account

If you are creating a new account:

  • After clicking on 'Create Account', enter your name and e-mail information and click 'Next'. Your e-mail information is very important.
  • Enter your institution and address information as appropriate, and then click 'Next.'
  • Enter a user ID and password of your choice (we recommend using your e-mail address as your user ID), and then select your area of expertise. Click 'Finish'.

Trouble Accessing an Existing Account

If you have an account, but have forgotten your log in details, go to ‘Password Help’ on the journal’s online submission system and enter your e- mail address. The system will send you an automatic user ID and a new temporary password.

Submitting Your Manuscript

  • After you have logged in, click the 'Submit a Manuscript' link in the menu bar. Enter data and answer questions as appropriate. You may copy and paste directly from your manuscript and you may upload your pre-prepared covering letter.
  • Click the 'Next' button on each screen to save your work and advance to the next screen.
  • You are required to upload your files.

-       Click on the 'Browse' button and locate the file on your computer.

-       Select the designation of each file in the drop-down menu next to the Browse button.

-       When you have selected all files you wish to upload, click the 'Upload Files' button.

  Main document, figures, “Supporting Information” and “Copyright Form” should be uploaded separately.
  A figure plate consisting of multiple figures should be uploaded as a single file and therefore the component figures should not be uploaded separately

  • Review your submission (in HTML and PDF format) before sending to the journal. Click the 'Submit' button when you are finished reviewing.



Online production tracking is available for your article through Wiley’s Author Services. Author Services enables authors to track their article, once it has been accepted, through the production process to publication online and in print. Authors can check the status of their articles online and choose to receive automated e-mails at key stages of production. The corresponding author will receive a unique link that enables them to register and have their article automatically added to the system. Please ensure that a complete e-mail address is provided when submitting the manuscript. Visit for more details on online production tracking and for a wealth of resources including FAQs and tips on article preparation, submission and more.



Proofs will be sent via email as an Acrobat PDF (Portable Document Format) file and should be returned within 3 days of receipt. Alterations to the text and figures (other than the essential correction of errors) are unacceptable at proof stage and authors may be charged for excessive alterations.


Acrobat Reader will be required in order to read the PDF. This software can be downloaded (free of charge) from the following website: This will enable the file to be opened, read on screen, and printed out in order for any corrections to be added. Further instructions will be sent with the proof.


Authors should supply an email address to which proofs can be emailed. Proofs will be faxed if no email address is available. If absent, authors should arrange for a colleague to access their email, retrieve the PDF proof and check and return them to the publisher on their behalf.



A minimum of 50 offprints will be provided upon request, at the author's expense. An Offprint Order Form outlining the cost of offprints will be sent to the corresponding author with the page proofs. Offprints will be provided only if a completed Offprint Order Form is returned to the publisher by the specified date.


Publication Fees

A publication fee of ¥12,000 is required per accepted manuscript if the first author is not a member of the Japanese Society of Phycology.


Regular Articles are limited to 8 printed pages and Research Notes are limited 4 printed pages. For manuscripts exceeding this limit, a page charge of ¥18,000 per additional printed page is levied to the author by the Japanese Society of Phycology. Approximately three typed pages will reduce to about one printed page. Figures and Tables are counted towards the page limit.


Authors should wait until they receive an invoice from the Japanese Society of Phycology before sending any fees. This procedure notwithstanding, no paper will be rejected or given extraordinary treatment on any basis other than its scientific merit.


Membership in The Japanese Society of Phycology is open to individuals and groups who have interest in phycology. Please visit the Society's website to get further information.


Early View Service

Phycological Research is covered by Wiley’s Early View service. Early View articles are complete full-text articles published online in advance of their publication in a printed issue. Articles are therefore available as soon as they are ready, rather than having to wait for the next scheduled print issue. Early View articles are complete and final. They have been fully reviewed, revised and edited for publication, and the authors' final corrections have been incorporated. Because they are in final form, no changes can be made after online publication. The nature of Early View articles means that they do not yet have volume, issue or page numbers, so Early View articles cannot be cited in the traditional way. They are therefore given a Digital Object Identifier (DOI), which allows the article to be cited and tracked before it is allocated to an issue. After print publication, the DOI remains valid and can continue to be used to cite and access the article. More information about DOIs can be found at

Address for Editorial Correspondence

Dr Mitsunobu Kamiya, Editor-in-Chief

Faculty of Marine Bioscience

Fukui Prefectural University

1 1 Gakuencho, Obama, Fukui

917 0003 Japan



Effective from January 2014