European Journal of Neuroscience

Cover image for Vol. 43 Issue 3

Edited By: John Foxe and Paul Bolam

Impact Factor: 3.181

ISI Journal Citation Reports © Ranking: 2014: 108/252 (Neurosciences)

Online ISSN: 1460-9568

Author Guidelines

Good reasons to publish in EJN

  • High standard, rigorous and fair peer-review
  • Rapid publication
  • Publishing is free and color figures appear in the online edition free of charge
  • EJN reaches a very wide audience and its readership continues to grow
  • Authors can choose to publish their accepted manuscripts as open access papers
  • EJN is the Journal of FENS
  • EJN is free to all the members of FENS societies and SfN
  • Publication proceeds fund the educational activities of FENS

The Editorial Office of EJN can be contacted at; Emma Boxer Tel. 441865476234

Click here to download the Author Guidelines as a PDF

1. General | 2. Types of manuscripts | 3. Manuscript Submission requirements | 4. Online submission | 5. Publication

1. General

1. General
1.1. Editorial Policy
1.2. Permissions
1.3. Competing interests
1.4. Funding
1.5. Sharing of datasets
1.6. Manuscript referral to Brain and Behavior

EJN is an international journal that publishes original research articles and reviews in the broad fields of developmental, molecular, cellular, systems, behavioral, and cognitive neurosciences. EJN aims to advance our understanding of organization and function of the nervous system in health and disease, thereby improving the diagnosis and treatment of neuropsychiatric and neurodegenerative disorders. Manuscripts submitted to EJN should describe novel results generated by experiments that were guided by clearly defined aims or hypotheses. Highly specialized articles with little scientific evidence, which provide only limited new insight into the organization and function of the nervous system, or insight that is too specialized for a generalist neuroscience journal may be considered outside the scope of EJN. EJN does not publish case reports.
EJN is a hybrid journal. Authors can opt to publish their accepted manuscripts as “locked", or open access papers (please see OnlineOpen).

EJN is a member of the Neuroscience Peer Review Consortium (NPRC), an alliance of neuroscience journals that have agreed to share manuscript reviews at the author’s request. The NPRC system allows fast-track reviewing of articles submitted together with prior reviews and appropriate revisions. EJN encourages authors of manuscripts that are of good quality but were rejected because of low priority rating to make use of this track when submitting to EJN. For more information, please visit

1.1 Editorial Policy
All studies should abide by the ethical standards as described in the Society for Neuroscience guidelines on Responsible Conduct Regarding Scientific Communication. EJN follows the Code of Conduct of the Committee on Publication Ethics (COPE) and handles cases of research and publication misconduct accordingly (

As per recommendation of ICMJE (, authorship at EJN is based on the following four criteria: "i) Substantial contributions to the conception or design of the work; or the acquisition, analysis, or interpretation of data for the work; ii) Drafting the work or revising it critically for important intellectual content; iii) Final approval of the version to be published and iv) Agreement to be accountable for all aspects of the work in ensuring that questions related to the accuracy or integrity of any part of the work are appropriately investigated and resolved […] All those designated as authors should meet all 4 criteria. And all who meeting all 4 criteria should be identified as authors. Those who do not meet all four criteria should be acknowledged […] All individuals who meet the first criterion should have the opportunity to participate in the review, drafting, and final approval of the manuscript. […] It is the collective responsibility of the authors, not the journal to which the work is submitted, to determine that all people named as authors meet all four criteria; it is not the role of journal editors to determine who qualifies or does not qualify for authorship or to arbitrate authorship conflicts. If agreement cannot be reached about who qualifies for authorship, the institution(s) where the work was performed, not the journal editor, should be asked to investigate." Any changes to the order of the authors, or the removal or addition of authors, needs to be approved by every author.

Whenever an author uses another person's words or ideas (in text, tables or figures), the author must cite the other person's work. If the author wants to use the text of another person word per word, this text should be in quotations and limited to a couple of sentences; the source should be referenced in the bibliography.

Submission of a manuscript to EJN is based on the understanding that the work described has not been published before, that is not under consideration for publication elsewhere, that its publication in the present form has been explicitly approved by all authors and by the responsible authorities in the institutions where the work was carried out, and that, if accepted, it will not be published elsewhere in the same form, in any language, without the consent of Wiley Blackwell. Anything published other than an abstract of less than 500 words is considered as prior publication. If authors would like to relate to such own previous publication, they should reference it in the bibliography in the same way they would refer to findings from a different research group. This policy applies to text, figures and parts of figures. Manuscripts that merely consist in expanded versions of work published elsewhere are not acceptable for publication in EJN. When submitting a manuscript, authors must provide (as supplementary document files) copies of any material they have published in the last two years (or which are in the process of publication) that overlaps the content of the manuscript they are submitting.

The CrossCheck plagiarism detection software is used to check submitted manuscripts for all forms of plagiarism, including self-plagiarism.

Submitted manuscripts are assigned to a section of the journal and an Associate Editor who is responsible for its evaluation. The Editor-in-Chief’s decision regarding publication is based on the recommendation of the Associate Editor and reports of reviewers,.

1.2 Permissions
If all or part of previously published illustrations are to be used, permission must be obtained from the copyright holder concerned. Wiley authors are free to re-use their own article or figures from their article in another publication providing they are author or co-author of the new publication. Address email queries to:

1.3 Competing interests
EJN requires that all authors disclose any potential sources of conflict of interest. Any interest or relationship, financial or otherwise, that might be perceived as influencing an author's objectivity is considered a potential source of conflict of interest. These must be disclosed when directly relevant or indirectly related to the work that the authors describe in their manuscript. Potential sources of conflict of interest include but are not limited to patent or stock ownership, membership of a company board of directors, membership of an advisory board or committee for a company, and consultancy for or receipt of speaker's fees from a company. The existence of a conflict of interest does not preclude publication in this journal. If authors have no conflict of interest to declare, they must also state this at submission. It is the responsibility of the corresponding author to review this policy with all authors and to collectively list in the cover letter to the Editor-in-Chief, in the manuscript under the Acknowledgement section and in the online submission system ALL pertinent commercial and other relationships.

1.4 Funding
All sources of funding should be declared in the Acknowledgements. If a private/commercial sponsor supported the research, authors are advised to describe the role of the study sponsor(s), if any, in study design; in the collection, analysis, and interpretation of data; in the writing of the report; and in the decision to submit the paper for publication. If the funding source had no such involvement, this should be stated.

1.5 Sharing of datasets
Before submission, data that cannot be published in the journal, such as large-scale datasets, must be deposited in one of the appropriate public repositories with private access that allows reviewers/Editors confidential, read-only access to data before manuscript acceptance. The accession codes must be provided in the materials and methods of the manuscripts; and the username and password must be indicated in the cover letter. Upon acceptance of the manuscript, records must be publically released (it is the authors’ responsibility to coordinate the date of release). Datasets for which there is no public database, should be submitted as Supporting document files.

1.6 Manuscript referral to Brain and Behavior
For manuscripts that are not accepted for publication in EJN, the authors can elect to have their manuscript transferred for consideration to Wiley's Open Access Journal, Brain and Behavior. The transfer will occur on-line and guarantee the anonymity of the peer-review process. It will not require reformatting or rewriting the manuscript at this stage. Brain and Behavior will render an editorial decision within a short time after the transfer and, if accepted, publish the manuscript typically within 15 days of acceptance. Brain and Behavior is an Open Access journal and article publication fees apply. For more information please go to

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2. Types of manuscripts

Research Reports
The major part of each Issue of EJN is devoted to Research Reports. These must describe significant and original scientific data from experimental or theoretical neuroscience studies and provide sufficiently detailed information in the Materials and method section for the observations to be critically evaluated and, if necessary, repeated. Research Reports do not normally exceed 12 printed pages in EJN, but the length of a manuscript will not, in itself, preclude publication. However EJN does not publish preliminary studies. In general, allow 1,350 words - including references and legends- per printed page and subtract 600 words for each table and figure.

Short Communications
Short Communications should report a significant advance in the field and should represent a ‘complete’ study rather than preliminary observations. They should contain a maximum of 3500 words, up to 4 Figures and 2 Tables and should be organized as described for Research Reports. The total word count must be indicated on the title page. We strongly discourage the use of supplementary material. Manuscripts not conforming to these guidelines may be returned to the authors or re-classified as a standard Research Report.

Review Articles
Review articles in EJN are full-length articles on topics of particular current interest. Extensive overviews of the literature are discouraged. Rather, review articles should aim to critically evaluate a timely issue of interest for the broad readership of EJN. While their length is not limited, the writing should be concise and precise. Illustrations, which effectively convey key topics of the review, are encouraged. Proposals for Review articles are welcomed by the Editors-in-Chief and the Associate Editors.

Technical Spotlight Articles
Technical Spotlight articles are brief reviews (total ~ 5000 Words) that evaluate and discuss novel techniques as well as controversial research approaches, or issues of validity concerning a major methodology or research avenue. Proposals for Technical Spotlight articles are welcomed by the Editors-in-Chief and the Associate Editors.

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3. Manuscript Submission requirements

3. Manuscript Submission requirements
3.1. Text formatting and English
3.2. Layout and file requirements
3.2.1. Original Submissions
3.2.2. Revisions and Resubmissions
3.2.3. NPRC submissions

3.2.4. Transferable Peer Review pilot
3.3. Title Page
3.4. Abstract
3.5. Introduction
3.6. Materials and methods
3.6.1. Ethical standards
3.6.2. Experimental animals
3.6.3. Suppliers
3.6.4. Reagents
3.6.5. Microscopy
3.6.6. Gels and Blots
3.6.7. Handling of image data
3.6.8. Statistical methods

3.7. Results and statistical analyses
3.8. Discussion
3.9. Acknowledgements
3.10. Abbreviations
3.11. References
3.11.1. Reference list
3.12. Tables
3.13. Figure legends
3.14. Figures
3.15. Graphical Abstracts
3.16. Supporting Information

Manuscripts should be prepared according to the guidelines below and submitted online at

3.1 Text formatting and English
The text of your manuscript should be prepared throughout in a standard font (Times/Times New Roman for text, and Helvetica/Arial and Symbol for figures). The manuscript should be formatted in double spacing and the lines should NOT be numbered. Please do not use a font size smaller than 12 points. Headings and sub-headings should not end with a period ".".

We recommend to authors for whom English is a second language to have their manuscript proofread by native English-speaking scientists. A list of Wiley Editing Services can be found at All services are paid for and arranged by the author, and use of one of these services does not guarantee acceptance or preference for publication.

3.2 Layout and file requirements
When uploading your files, please use simple filenames. Filenames must not include any spaces or special characters (do not use underscores). The submission files should not exceed 250MB in total. If you have trouble loading large files, please contact the editorial office at for assistance.

For manuscripts that contain complex equations or other mathematical content, it is possible to submit LaTeX files. If you do, please also upload the PostScript or PDF version of your compiled file.

3.2.1 Original Submissions
The manuscript should be uploaded either in the format listed under Revisions (see below), or as a single PDF file (containing the main text document, tables and figures). In the latter case, figures must be at the end of the PDF document and their number must be indicated on the same page as the figures. Your cover letter should not be part of the PDF.

The sections of a Research Report should be in the following order: Title page, Abstract, Introduction, Materials and methods, Results, Discussion, Acknowledgements, Abbreviations, References, Tables, Figure Legends and Figures. Review articles and Technical Spotlight articles are not required to follow this outline.

3.2.2 Revisions and Resubmissions
To submit a revised manuscript, you must submit a word document file (saved as a .doc or .docx file) and high resolution figures separately (for figure requirements, please see 3.13 Figures section). You may NOT submit PDF files for revised manuscripts, except for the supporting document and the reply to the reviewers. A detailed point-by-point response letter to the reviews, describing the changes you have made in the text should be uploaded as a separate Word or PDF file using the file designation "Authors' Responses to Reviewers". For Revisions ONLY changes to the text should be emboldened or underlined to assist reviewers with re-evaluation (PLEASE DO NOT USE TRACK CHANGES). For resubmissions, changes to the text should not be outlined. All supporting information, including supp. figure legends, should be merged and uploaded as a separate single PDF file, except video files that should be uploaded with the file designation “Media files”.

Files should appear in the following order: Reply to Reviewers, Title page, Abstract, Introduction, Materials and methods, Results, Discussion, Acknowledgements, Abbreviations, References, Tables, Figure Legends and Figures, Graphical Abstract Text, Graphical Abstract Figure.

3.2.3 NPRC submissions
For manuscripts submitted through the NPRC track, please ask the editorial office of the previous journal to transfer the reviews along with the name of the reviewers to In addition to submitting your manuscript as an original submission, please submit a detailed point-by-point response letter to the previous reviews by uploading a Word or PDF file with the file designation "Authors' responses to reviewers".

3.2.4 Transferable Peer Review Pilot
European Journal of Neuroscience is participating in Wiley's pilot of transferable peer. Authors of original research articles that are rejected from the journal with completed reviews will be invited to transfer the manuscript, accompanied by the previous reviews, to any of the other journals participating in the pilot. Authors will have the opportunity to revise their manuscript according to the reviewers’ comments prior to transfer if they wish to do so. A list of participating journals and more information about the pilot can be found here. We believe that this system of preserving original peer review for the next journal's use will decrease repetitious review, save authors, reviewers and editors valuable time, and significantly increase speed to publication for many papers.

Please note that this transferable peer review scheme is separate from the NPRC transfer system.

3.3 Title Page
The Title page must include:

  • The proposed Journal section
  • A clear and concise title
  • The author's first name(s) and surnames
    Note: Authors who normally write their names in non-Latin characters may include both a transliterated version of their names, followed in parenthesis by their names in their native writing system. Any non-Latin languages that can be represented in Unicode characters will be accepted.
  • The address(es) from which the work originated
  • The name, institutional affiliation, address, fax number and e-mail address of the person who will deal with correspondence, including proofs (which are sent electronically)
  • A running title not in excess of 50 characters and spaces
  • The total number of pages, figures, tables and equations
  • The total number of words in: (i) the whole manuscript; (ii) the Abstract; and (iii) the Introduction.
  • A list of four or five keywords not appearing in the title, preceded by “Keywords”.

If the species used does not appear in the title, it should be included in the keywords and be mentioned in the first few lines of the abstract.

3.4 Abstract
The Abstract should be written in complete sentences without headings and should provide a summary not exceeding 250 words, in a form comprehensible to any neuroscientist and suitable for publication without the full article text. The aim of the study and working hypothesis should be stated in the Abstract, and the animal species used indicated, if it is not given in the title. The Abstract should provide a concluding statement highlighting the significance and impact of the results for the neurosciences. If references must be cited in the Abstract they must include the author(s), journal title, volume number, page span, and year. The use of abbreviations and statistical results should be avoided in the abstract.

3.5 Introduction
The main part of a Research Report should start with a brief Introduction not exceeding 500 words, which outlines the historical or logical origins of the study and clearly states the aim of the study and/or hypothesis to be tested, without repeating the Abstract or summarizing the results.

3.6 Materials and methods
The Materials and methods section should provide a sufficiently detailed description of the methods to allow a researcher to reproduce your work. Important methodological aspects of your work, such as generation of mutant animals or the method of sample preparation, should be described, even if such descriptions can also be found in prior publications.

3.6.1 Ethical standards
Studies using human or animal subjects should include an explicit statement identifying the Institution or Review Committee which approved the study. Editors reserve the right to reject papers if there is doubt whether appropriate procedures were followed.

(i) Studies involving human subjects
When human subjects are used, manuscripts must be accompanied by a statement that the experiments were undertaken with the understanding and written consent of each subject, and that the study conforms with World Medical Association Declaration of Helsinki published on the website of the Journal of American Medical Association in 2013. In addition, the name of institutional review board (or appropriate committee of the institution) that approved the study should be given and their approval should be explicitly stated.

(ii) Studies involving experimental animals
The Materials and methods section must briefly but explicitly state measures which were taken to minimize pain or discomfort, e.g. type and dose of anesthetic used. Experiments should be carried out in accordance with the Council Directive 2010/63EU of the European Parliament and the Council of 22 September 2010 on the protection of animals used for scientific purposes or with the Guidelines laid down by the NIH in the US regarding the care and use of animals for experimental procedures. In addition, the name of the Animal Use and Care Committee or Institution that approved the study should also be given and their approval should be explicitly stated.

3.6.2 Experimental animals
When experimental animals are used, specify species, strain, sex, age, supplier, and numbers of animals used in total and for individual experimental conditions. The species should be identified in the Title or Abstract.

If genetically modified mice were used, the Standards for the publication of mouse mutant studies (see Crusio et al., Genes, Brain and Behavior (2009) 8:1–4) should be followed. Provide detailed and full strain and substrain information and use the correct nomenclature for identifying strains (see Crusio et al. 2009).

3.6.3 Suppliers
Suppliers of materials should be named and, with the exception of well-known suppliers, such as Sigma, Kodak and Zeiss, their location (city, state, country) included.

3.6.4 Reagents
The origin and specificity of reagents, notably antibodies, oligonucleotides, cDNA constructs, need to be documented, including the nucleic acid sequence or peptide sequence used. Nucleic acid probes should be fully sequenced when applicable. Control experiments that were conducted to ensure the specificity of the method (immunohistochemistry, immunoprecipitation, Western blotting, in situ hybridization) should be described, along with key references to previous work with this reagent. For antibodies, this documentation must include a precise description of the antigen, the nature of the antibody (species, purification), the supplier, catalogue number, and specificity tests performed (e.g., Western blot, immunoprecipitation, validation with knockout tissue, etc). For reports involving immunohistochemical methods, please consult the criteria and recommendations discussed in Fritschy JM (2008) EJN 28:2365-2370: and the editors' tips in the EJN News at

3.6.5 Microscopy
Information necessary to evaluate the resolution of images from microscopy (type of microscope, numerical aperture of the lens) must be provided. Microscopic images must include calibration bars (statements about original magnification are insufficient, as final published images can be any size). For confocal laser scanning microscopy, the voxel size and the number of confocal planes included in an image must be indicated. For quantification of digital parameters (signal intensity, size or number of objects), detailed information about the procedure, notably criteria for threshold determination, normalization, and calibration (if applicable) should be provided.

3.6.6 Gels and Blots
Gels and blots should be presented with molecular weights noted. Cropping of images for clarity and conciseness purposes is acceptable but at least several band widths should be kept above and below the cropped band and important bands must be shown. Merging images from different experiments is not acceptable. If splicing of data from a single experiment is necessary to reorder the samples, this should be clearly indicated on the figure and in the figure legend. The method of normalization of results must be explicitly stated in the figure legends.

3.6.7 Handling of image data
EJN follows the guidelines set by the Rockefeller University Press and endorsed by the Council of Science Editors

  • No specific feature within an image may be enhanced, obscured, moved, removed, or introduced.
  • Adjustments of brightness, contrast, or color balance are acceptable if they are applied to the whole image and as long as they do not obscure, eliminate, or misrepresent any information present in the original.
  • The grouping of images from different parts of the same gel, or from different gels, fields, or exposures must be made explicit by the arrangement of the figure (e.g., dividing lines) and in the text of the figure legend.
  • If the original data cannot be produced by an author when asked to provide it, the acceptance of the manuscript may be revoked.

3.6.8 Statistical methods
A complete description of statistical methods is required. The recommendations described in the EJN Editorial on Reporting Statistical Methods and Results (EJN 2008, 28, 2363-2364) are a component of these Author Guidelines.

3.7 Results and statistical analyses
The observations should be presented with minimal reference to earlier literature or to possible interpretations. The main statistical results should be reported in the Results section. The description of the statistical results should include the proper statistical term (such as the F statistic) as well as the degrees of freedom and the P value. The description of statistical results in the figure legends should be limited to important post hoc comparisons. The recommendations described in the EJN Editorial on Reporting Statistical Methods and Results (EJN 2008, 28, 2363-2364; are a component of these Author Guidelines and should be followed in detail.

3.8 Discussion
The Discussion should begin by summarizing the major findings, while avoiding repetition of statements provided in the Abstract or the Results section. For clarity, the use of subheadings is recommended.

3.9 Acknowledgements
A short statement about grant and other financial support should be given, along with a list of contributions from collaborators who are not co-authors (it is implicit that they agree with this mention), and a declaration of competing interests. See above under Editorial Policies for additional items to be addressed in the Acknowledgements.

3.10 Abbreviations
Abbreviations should be kept to an absolute minimum. Abbreviations save relatively little space but greatly diminish the readability of a manuscript. In general, abbreviations should not appear in the Abstract, and sentences that contain more than one abbreviation merit careful review. Abbreviations must be used more than four times in an article, and consistently, before their use can be permitted. The word must always be written out in full when first used and the proposed abbreviation given in parentheses. A list of all abbreviations used in the text and their meanings must be provided (in alphabetic order).

3.11 References
All references must be listed, and all listed references must be cited at least once in the main text (particular care is needed to check this after any modification or revision of the text). Citations in the text should be given in parenthesis, e.g. (Blanc & White, 1985; Weiss et al., 1986), except when the author’s name is part of a sentence, e.g. “White (1985) reported that … “. Where there are two authors, both should be named, but with three or more, only the first author’s name plus ‘et al´ should be given. Note that when more than one paper refers to the same first authors in the same year, a, b, c suffixes must be used in the text and reference list to avoid ambiguities, written as (Weiss et al., 1986a; White, 1986a,c).

3.11.1 Reference list
The use of a tool such as EndNote, Reference Manager, or Bookends is recommended for reference management and formatting. EndNote reference styles can be searched for at, and Reference Manager reference styles can be searched for at

There should be a list of references in alphabetic order according to the name of the first author and then chronologically where several papers by the same author are cited. All authors should be included in each reference appearing in the Reference List; these should not be abbreviated to et al. Only published and ‘In Press’, i.e. accepted for publication in a specific journal or book, references should be included in the reference list. No characters (e.g. numbers or tabs) should appear before the first author's name.

(i) Journal Articles
The titles of journals should be abbreviated in accordance with the World List of Scientific Periodicals, 4th edition. Single-word titles, e.g. Psychoneuroendocrinology, should not be abbreviated.
Journal articles should be formatted as follows:

Lancaster, B. & Boxall, A. R. (1998) Tyrosine kinases and synaptic transmission. Eur. J. Neurosci., 10, 2-7.
Note the emboldening of the Volume number.

(ii) Books and Symposia
Books should be listed as follows:

Katz, D.I. (1997) Traumatic brain injury. In Mills, V.M., Cassidy, J.W. & Katz, D.I. (eds), Neurologic Rehabilitation. A Guide to Diagnosis, Prognosis and Treatment Planning. Blackwell Science, Oxford, pp. 105-143.

Symposia contributions should take a similar format to books, including the place and date of the meeting, the name and location of the organiser/publisher, both overall and contribution titles, all authors and Editors names, and page number(s):
Fuss, S.H., Çelik, A. & Korsching, S.I. (2001) Levels of olfactory information processing in the zebrafish olfactory bulb. In Elsner, N. & Kreuzberg, G.W. (eds), Göttingen Neurobiology Report 2001. Proceedings of the 4th Meeting of the German Neuroscience Society 2001, Vol. II, 28th Göttingen Neurobiology Conference. Georg Thieme Verlag, Stuttgart, p. 474.

3.12 Tables
Tables must be supplied as editable text and not as embedded figures/objects (except for original submissions, where it is possible to submit a single PDF file). Tables should be numbered, and have a bold title and appear in the text following the references. All columns should have a heading.

3.13 Figure legends
Figure legends should be listed as part of the main manuscript text file at the end of the document. These should explain each figure as fully as possible, referring the reader to the text only on rare occasions in order to avoid repeating in the legends material that must be included in the text. All abbreviations used in the figures should be explained in each legend when there are less than five abbreviations used; for legends with large numbers of abbreviations, the authors should include these in the abbreviations list (see above). Scale bars are required for all photographs. Main statistical findings should be described in the Result section. The legends may describe the results of important post-hoc comparisons (usually also symbolized in graphs depicting data).

3.14 Figures
All figures and tables should be uploaded separately and must be labeled with a figure number (except for initial submissions, where it is possible to submit a single PDF file). Before uploading figure files, all white/blank spaces around the figure should be cropped out. To ensure accurate conversion of characters, standard fonts such as Times, Times New Roman, Ariel or Helvetica should be used for preparing the text and figures. Symbol font should be used to convert Greek characters and other non-standard characters such as the degree symbol.

For production, figures need to be of a high resolution and saved as .tiff files (300 d.p.i. for colour/half tone and 800 d.p.i. for line work (black line art, white background), or saved as .eps or .doc files. Photoshop or IrfanView (free download on Web) may be used to produce high resolution figures. In their desired final size, figures need to fit the width of a single column of text, i.e. 88 mm wide, or where necessary they should be 184 mm in maximum width. The final size of the printed figure cannot exceed 230 x 184 mm including legends, and where a reduction is required this should be indicated on the figures. Any lettering should be no less than 2 mm in height in the printed figure and should be in proportion to the overall dimensions of the figure. Where several figures are mounted together they should be squared accurately and separated by about 5 mm. All of the figures in such a group should have approximately the same contrast values.

The initial upload of lower resolution images is sufficient (.jpg files allowed; 2 MB maximally); upon revision / acceptance high resolution TIFF images are requested.

Avoid bar or line graphs with only two data points, such results should be described in the text only. There is no limit to the number of figures per manuscript, but authors should use them sparingly, making sure that each figure is relevant to the text and that the figures are presented and numbered in the order in which they are mentioned in the text. When there are large numbers of figures, layout problems are less likely to occur if the figures are of similar shapes and sizes. The submission files should not exceed 250MB in total. If you have trouble loading large files, please contact the editorial office at for assistance.

See above the subsection Handling of image data for instructions about image manipulation.
Please refer to for detailed guidelines on electronic artwork.

Color figures
Authors can elect to have color illustrations only in the online version of their published manuscript while having them reproduced black-and-white in the printed version, free of charge. In this case, (a) a color version as well as a black-and-white version of the figure should be uploaded, and (b) the figure legend should not refer to color as it will be used for both print and online versions.

If authors elect to have color figures published in the printed journal, color reproduction costs are charged per printed page of color. The cost per printed page with colour is £450 VAT. These charges are waived for all Review articles, and on request for authors from former Eastern Bloc countries publishing Research Reports. Color figures that are sequentially numbered can be placed on the same page to reduce costs. Avoiding a mix of color panels with black-and-white ones can reduce costs, improve reproduction and allow more space for the color.

Following acceptance, a signed copy of the completed Color Work Agreement Form must be sent to Customer Services before color work can be processed. The Journal allows authors to publish figures in color free of charge in the online edition if requested on submission. Authors must then indicate they wish to publish in black-and-white in the print edition and in colour in the online edition. In this case, (a) please supply a black-and-white as well as color version of the figure and (b) the figure legend should not refer to color as it will be used for both print and online editions.

3.15 Graphical Abstracts
Graphical abstracts are aimed at promoting articles. They appear in the online tables of contents of EJN and in the list of Early View articles.

For all revised and resubmitted manuscripts, please upload the following files in step 6 of the submission process:

  • a Word document file with the file designation "Graphical Abstract Text". The graphical abstract text should consist of 2-3 sentences (maximum 500 characters with space) highlighting the major findings of the article and their impact for the neurosciences.
  • a figure file with the file designation "Graphical Abstract Figure". The figure should be representative of your manuscript and should contain no caption, heading or outline. Labels on the figure may be removed when appropriate. The figure needs to be of a high resolution and saved as a .tiff file (300 d.p.i. for color/half tone and 800 d.p.i. for line work), or saved as .eps or .doc files. Before uploading the file, all white/blank spaces around the figure should be cropped out.

3.16 Supporting Information
The use of Supporting Information should be limited to non-essential items, such as documentation of experiments controlling the specificity of reagents, etc. Authors need to justify in the cover letter the need for supporting information in their manuscript.

Supporting Information will not be published in the print edition of the journal but will be included in the online edition, or as a link to the information given. It has to be submitted for review purposes. Supporting information should appear at the end of the manuscript, i.e. following the figures.

Supporting information will be published as submitted and will not be edited, thus authors should make sure the data is clearly presented and that it conforms with the style of the rest of the paper. Supporting information should be merged and uploaded as a single PDF file, except for video files that should be uploaded with the file designation “Media files”.

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4. Online submission

Authors should submit their manuscripts online at

There are no submission fees or page charges to submit a manuscript to EJN for black-and-white print. Authors can also request to get their figures published in color in the online journal for free. Alternatively, authors can request to get their figures published in color in the printed journal, in which case color-printing costs will be charged per printed page of color.

During the submission process the following must be provided:

  • The names, valid emails and institutions of all of the authors included in the paper
  • The name of the most appropriate journal section and Section Editor should be selected from the list provided. Please choose a Section Editor in whose area of expertise your manuscript falls – To view the list of Section Editors with their areas of expertise and journal sections, please refer to
  • All manuscripts files:
    • Main text, including Title page, Acknowledgements, Abbreviations and Figure Legends
    • Tables and figures
    • For original submissions ONLY, you may submit a single PDF files containing the main text document, tables and figures (see above for details)
    • If the manuscript is a NPRC submission, a revision or resubmission, please upload in step 6 of the submission process a point-by-point response letter, describing the changes you have made in the text. Please upload this response letter as a separate Word or PDF file, using the file designation "Authors' Responses to Reviewers".
  • If the manuscript is being resubmitted, you will need to provide the original manuscript ID

Cover illustrations
If you have any color illustrations, either appearing in your paper or relating to it, that would make good images for the front cover, please upload them as JPG or TIFF files during the file upload step of the submission process, with the file designation "Cover Image Suggestion".

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5. Publication

5. Publication
5.1. License Agreements
5.1.1. Authors funded by NIH
5.1.2. Authors funded by the Wellcome Trust and Research Councils UK (RCUK)
5.1.3. Other funder policies
5.1.4. OnlineOpen

5.2. Self archiving
5.3. Accepted Articles
5.4. Early View Articles
5.5. Offprints

When a manuscript is accepted, authors will be advised that their accepted manuscript has been electronically sent from the Editorial Office to the Production Editors of the Publishers, Wiley Blackwell. The Production Editor can be contacted at Authors will be notified to send a completed and signed copy of the Color Work Agreement Form if color figures are required, to Customer Services before their accepted manuscript will be processed for publication. Also it is necessary to send evidence of permission to use any previously published material.

5.1 License Agreements
EJN is a hybrid journal. Authors can opt to publish their accepted manuscripts as “locked”, or “open access” papers (OnlineOpen). Upon acceptance of a manuscript, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper. If the OnlineOpen option is not selected the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs:

5.1.1 Authors funded by NIH
The NIH mandates grantees to deposit their peer-reviewed author manuscripts in PubMed Central, to be made publicly available within 12 months of "general" publication. The NIH mandate applies to all articles based on research that has been wholly or partially funded by the NIH and that are accepted for publication on or after April 7, 2008. In order to help authors comply with the NIH mandate, for papers accepted for publication in EJN, Wiley Blackwell will post the accepted manuscript (incorporating all changes made during peer-review, but prior to the publisher's copyediting and stylistic edits, online and print formatting, citation and other linking, deposit in abstracting and indexing services to enable high-level discovery, and the addition of bibliographic and other metadata) of NIH grant-holders to PubMed Central at the point of acceptance by the journal. This version will then be made publicly available in PubMed Central 12 months after publication. Following the deposit authors will receive further communications from the NIH with respect to the submission. For further information, see here.

Alternatively, if authors wish to make their FINAL published article open access and without a 12 month embargo, they can choose to publish via the OnlineOpen service. See below for further details.

5.1.2 Authors funded by the Wellcome Trust and Research Councils UK (RCUK) or the Austrian Science Fund (FWF)
If you select the OnlineOpen option and your research is funded by certain funders [e.g. The Wellcome Trust and members of the Research Councils UK (RCUK) or the Austrian Science Fund (FWF)] you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with your Funder requirements. For more information on this policy and the Journal's compliant self-archiving policy please visit:

5.1.3 Other funder policies
For information for HHMI grantees and other funder policies, please visit

5.1.4 OnlineOpen
OnlineOpen is available to authors of articles who wish to make their article open access. With OnlineOpen the author, their funding agency, or institution pays a fee to ensure that the article is made available to non-subscribers upon publication via Wiley Online Library, as well as deposited in PubMed Central and PMC mirror sites. In addition to publication online via Wiley Online Library, authors of OnlineOpen articles are permitted to post the final, published PDF of their article on a website, institutional repository, or other free public server, immediately on publication. For full details please see

You will not be requested to inform the editorial office that you intend to publish your article as OnlineOpen before acceptance. OnlineOpen articles go through the same peer-review process as any other article. After acceptance, if you want your article to be open access please choose the appropriate licence agreement when you log in to Wiley’s Author Services system. Click on ‘Make my article OnlineOpen’ and choose the appropriate license by clicking on ‘Sign license agreement now’ when you log in to Wiley’s Author Services system.

If the OnlineOpen option is selected the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):

  • Creative Commons Attribution License OAA
  • Creative Commons Attribution Non-Commercial License OAA
  • Creative Commons Attribution Non-Commercial -NoDerivs License OAA
To preview the terms and conditions of these open access agreements please visit the Copyright FAQs and

Government employees in both the US and the UK need to complete the Author Warranty sections, although copyright in such cases does not need to be assigned. After submission of the form, authors will retain the right to publish their paper in various media/circumstances. Please see the form for further details and email queries to:

5.2 Self archiving
Authors are permitted to deposit the accepted version of their article (pre-copy editing version) in personal, subject, or institutional repositories 12 months after publication. In addition, authors of OnlineOpen articles are permitted to post the final, published PDF of their article on a website, institutional repository or other free public server, immediately on publication.

5.3 Accepted Articles
'Accepted Articles' have been accepted for publication and undergone full peer review but have not been through the copyediting, typesetting, pagination and proofreading process. Accepted Articles are published online a few days after final acceptance, appear in PDF format only, are given a Digital Object Identifier (DOI), which allows them to be cited and tracked, and are indexed by PubMed. Therefore the submitting author must carefully check the names and affiliations of all authors provided in the cover page of the manuscript, as it will not be possible to alter these once a paper is made available online in Accepted Article format.

5.4 Early View Articles
EJN articles are published online in advance of their publication in a printed issue through the Early View service (access Early View articles of EJN at Early View articles have been fully reviewed, revised and edited for publication, and the authors' final corrections have been incorporated. Because they are in final form, no changes can be made after online publication.

5.5 Offprints
Free access to the final PDF offprint or your article will be available via author services only. Please therefore sign up for author services if you would like to access your article PDF offprint and enjoy the many other benefits the service offers.

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