Acta Zoologica

Cover image for Vol. 97 Issue 1

Edited By: Lennart Olsson

Impact Factor: 1.258

ISI Journal Citation Reports © Ranking: 2014: 13/21 (Anatomy & Morphology); 63/154 (Zoology)

Online ISSN: 1463-6395

Author Guidelines


Submissions to Acta Zoologica are now made on-line using ScholarOne Manuscripts. To submit to the journal go to If this is the first time you have used the system you will be asked to register by clicking on ‘create an account’.  Full instructions on making your submission are provided.  You should receive an acknowledgement within a few minutes.  Thereafter, the system will keep you informed of the process of your submission through refereeing, any revisions that are required, and a final decision.

Manuscripts should be in English and written in a concise, straightforward style. Authors for whom English is a second language may choose to have their manuscript professionally edited before submission to improve the English. A list of independent suppliers of editing services can be found here. All services are paid for and arranged by the author, and use of one of these services does not guarantee acceptance or preference for publication.

The manuscript should present scientific findings which are essentially new and which have not been published or submitted for publication elsewhere. Review papers are also welcomed.

Papers of up to 24 printed pages (including illustrations, tables, references, etc.) are printed without page charge. Authors who wish to submit larger papers are advised to contact the editor in advance. All correspondence will normally be through the Editorial Office.

The manuscript should be typed double-spaced on one side of white A4 paper, with wide margins. All pages should be numbered. Original illustrations should not be sent until the manuscript has been accepted for publication. The form and content should be checked carefully to exclude the need for corrections in proof. Changes in the proof other than typographical errors will be charged to the authors.

Where a paper has joint authors, one author must accept responsibility for all correspondence.

During the submission process the following must be provided:

  • The names, emails and Institutions of all of the authors included in the paper.
  • The names, Institutions and emails of two suggested reviewers. Please do not suggest scientists based in your own institution or close collaborators elsewhere, as this will be viewed as a conflict. It is at the discretion of the Editors whether the suggested reviewers are approached for a review.
  • If the manuscript is a revision or re-submission, please upload a point-by-point response statement in the `Response to decision letter' field. Please do not upload this as a manuscript file. Major changes to the text should be emboldened or underlined to assist reviewers with re-evaluation.

Title Page
Title, name(s) of author(s), institution(s), full address(es), email address(es) and a condensed title of up to 45 characters to be used as the running head.

The names, emails and Institutions of all of the authors included in the paper

The names, Institutions and emails of two suggested reviewers. Please do not suggest scientists based in your own institution or close collaborators elsewhere, as this will be viewed as a conflict. It is at the discretion of the Editors whether the suggested reviewers are approached for a review.

If the manuscript is a revision or re-submission, please upload a point-by-point response statement in the `Response to decision letter' field. Please do not upload this as a manuscript file. Major changes to the text should be emboldened or underlined to assist reviewers with re-evaluation

Four grades of headings may be used:

A new line, full out left, initial capital letter main words, bold type; following text on new line not indented; [example Format]

B new line, full out left, initial capital letter first word only, italic; following text on new line not indented; [example Headings]

C new line, full out left, initial capital letter first word only, italic; text follows on same line after full point; [example Principal (A) headings]

D new line, indented, initial capital letter first word only, italic; text follows on same line after full point; [example Abstract]

Please indicate the grade of headings clearly in the left margin of the manuscript. B, C and D headings should be underlined to indicate italics.

Principal (A) headings
The following A headings should be used as and where appropriate.

Abstract. This should be an informative, condensed version of the important results and discussions and should not exceed 200 words. The abstract should be preceded by bibliographic details and followed by the name and full postal address of the author to whom correspondence should be addressed.

Lester. S. M. 1988. Ultrastructure of adult ... normani. - Acta Zoologica (Stockholm) 69: 95-109. [abstract]
Susan M. Lester. Department of., CA 95616, U.S.A. E-mail: xxx@xxx.xx.xx (or fax number if e-mail is not available).

Materials and Methods.


References in Articles
We recommend the use of a tool such as EndNote or Reference Manager for reference management and formatting.

EndNote reference styles can be searched for here:

Reference Manager reference styles can be searched for here:

A list of all the publications cited in the paper (and no others). The style of the references can be seen from the following examples.

Hatschek, B. 1877. Embryonalentwicklung und Knospung der Pedicellina echinata. - Zeitschrift für wissenschaftliche Zoologie 29: 502-549, pls 28-30.

Clark, R. B. 1980. Natur und Entstehung der metameren Segmentierung. - Zoologische Jahrbücher, Anatomie 103: 169-195.

Cloney, R. A. 1978. Ascidian metamorphosis: review and analysis. In F.-S. Chia and M. E. Rice (Eds): Settlement and Metamorphosis of Marine Invertebrate Larvae, pp. 255-282. Elsevier, New York.

Ruppert, E. E. 1997. Cephalochordata (Acrania). In Harrison, F. W. (Ed.): Microscopic Anatomy of Invertebrates, vol. 11, pp. 349-504. Wiley-Liss, New York.

Salvini-Plawen, L. V. and Mayr, E. 1977. On the evolution of photoreceptors and eyes. - Evolutionary Biology 10: 207-263.


Files should be formatted double-spaced with no hyphenation and automatic word wrap (no hard returns within paragraphs). Please type your text consistently, e.g. take care to distinguish between '1' (one) and l (Lower case L) and '0' (zero) and 'O' (capital O), etc.

Latin names of genera and species should be in italics or underlined to indicate italics. All other indications for the printer will be inserted by the editor.

Literature references in the text should have the following form:

'Hatschek (1877) described...' or 'The statement by Hatschek (1877, p. 506) 'Die Furchungselemente ... grösser' has been...' or 'As described earlier (Hatschek 1877; Reed and Cloney 1983a, b; Strathmann et al. 1972, fig. 2)...'

Footnotes should generally be avoided.

Reference to figures and tables in the manuscript should have the following form: 'As shown in Fig. 3 and Table 1...' or 'The cells have many cilia (Fig. 3)'.

Supporting Information

Supporting Information can be a useful way for an author to include important but ancillary information with the online version of an article. Examples of Supporting Information include additional tables, data sets, figures, movie files, audio clips, 3D structures, and other related nonessential multimedia files. Supporting Information should be cited within the article text, and a descriptive legend should be included. It is published as supplied by the author, and a proof is not made available prior to publication; for these reasons, authors should provide any Supporting Information in the desired final format.

For further information on recommended file types and requirements for submission, please visit:


Please send us digital versions of your figures if at all possible. Line illustrations or photographs should be supplied after the manuscript has been accepted. Line illustrations should be provided at 600 dots per inch (dpi) and photographs at 300 dots per inch in either TIFF or EPS format. Further information on these formats is available at  

All illustrations are to be numbered consecutively with arabic numerals, and each one must be referred to in the text. Photographs arranged in groups together should all form part of the same figure. For example, four photographs should be referred to as Fig. 1A, 1B, 1C and 1D rather than figures 1-4. The approximate position of illustrations and tables should be indicated in the margin of the text. All illustrations should be marked with the author's name. The legends should be self-explanatory and must always be included below the illustration and on the same page. Please note that your paper will go through production more quickly if the figures do not have to be relabelled or redrawn. Please ensure that electronic artwork is prepared such that, after reduction to fit across one or two columns or two-thirds width (80mm, 169mm or 110mm, respectively) as required, all lettering will be clear and easy to read, i.e. no labels should be too large or too small. Avoid using tints if possible; if they are essential to the understanding of the figure, try to make them coarse. No artwork should be incorporated into the text files. Original drawings or photographs should be supplied for reproduction.

The illustrations should preferably be submitted as photographic reproductions (not photocopies) at the size intended for printing. This will enable authors both to judge the suitability of the illustrations for printing and keep a standardised lettering, viz. 10 point letters for line drawings and 12-14 point letters for photographs.

Photographs should be provided as glossy prints with good contrast.

The magnifications of the illustrations should be indicated by scale bars.

Full instructions concerning the submission of artwork can be found on the journal’s Manuscript Central site:


Tables should be typed as text, using 'tabs' to align columns. The use of table editors should be avoided. Do not use graphics software to create tables.

Full instructions concerning the submission of tables can be found on the journal’s Manuscript Central site:

File Description Form
The submitted manuscript must be accompanied by the completed File Description Form. Until this is received, the manuscript cannot be put into production.

Acta Zoologica welcomes colour figures and photographs. All black and white figures are free in both the printed and online versions of an article. All colour figures on the online version and in the PDF offprints are free of charge. Colour figures in the printed version of an article are charged at a cost.

Therefore, please note that if there is colour artwork in your manuscript when it is accepted for publication, Wiley Blackwell requires you to complete and return a colour work agreement form before your paper can be published. This form can be downloaded as a PDF here. If you are unable to download the form, please contact the Production Editor.

Once completed, please return the form by mail or courier to the following address:

Customer Services (OPI)
John Wiley & Sons Ltd, European Distribution Centre
New Era Estate
Oldlands Way
Bognor Regis
West Sussex
PO22 9NQ

Full details are given on the form. Under exceptional circumstances, authors may request the charges to be waived. This must be done, with justification from the author, at the time of submission of the manuscript. If the author feels that inclusion of the figure(s) in colour is essential for interpretation of the results presented, then the author must provide justification to the Editors in writing at the time of submission. If authors wish to apply for funds to cover the costs of colour printing, the Editors will provide relevant support letters to funding bodies, indicating acceptance of the paper.

New Zoological Taxon
When publishing papers that describe a new zoological taxon, Acta Zoologica complies with the amendment to the International Commission on Zoological Nomenclature Code. Authors have to register any new species to Zoobank and obtain a Life Science Identifier (LSID) (e.g., XXXXXXX). These numbers must be provided together with the submitted manuscript. In the final published paper (Version of Record), the LSID will be listed under the new species name in the taxon section heading. Registration must be done as early as possible to avoid delay of publication upon acceptance of the manuscript.

Data archiving
Data that is integral to the paper must be made available in such a way as to enable readers to replicate, verify and build upon the conclusions published in the paper. Any restriction on the availability of this data must be disclosed at the time of submission.

Data may be included as part of the main article where practical. We recommend that data for which public repositories are widely used, and are accessible to all, should be deposited in such a repository prior to publication. The appropriate linking details and identifier(s) should then be included in the publication and where possible the repository, to facilitate linking between the journal article and the data. If such a repository does not exist, data should be included as supporting information to the published paper or authors should agree to make their data available upon reasonable request.

Proofs are now available through the web. The corresponding authors will be sent an email with full instructions from the typesetters. They should be returned to the Production Editor within 3 days of receipt. Where this is not possible authors should email the Production Editor ( to advise of the delay. Proofs can be faxed back to the Production Editor on +65 6643 8599. The return address for proof corrections is as above. Please do not use Adobe Acrobat software options to electronically edit the proof, as doing so may delay the publication of your paper. Major alterations to the text and illustrations are only accepted when absolutely necessary; the additional cost may be charged to the author.

Free access to the final PDF offprint or your article will be available via Author Services only. Please therefore sign up for Author Services if you would like to access your article PDF offprint and enjoy the many other benefits the service offers.

Additional paper offprints may be ordered online. Please click on the following link, fill in the necessary details and ensure that you type information in all of the required fields.

If you have queries about offprints please email

Early View
Acta Zoologica is covered by Wiley Blackwell’s Early View service.
Early View articles are complete full-text articles published online in advance of their publication in a printed issue. Articles are therefore available as soon as they are ready, rather than having to wait for the next scheduled print issue. Early View articles are complete and final. They have been fully reviewed, revised and edited for publication, and the authors’ final corrections have been incorporated. Because they are in final form, no changes can be made after online publication. The nature of Early View articles means that they do not yet have volume, issue or page numbers, so Early View articles cannot be cited in the traditional way. They are therefore given a Digital Object Identifier (DOI), which allows the article to be cited and tracked before it is allocated to an issue. After print publication, the DOI remains valid and can continue to be used to cite and access the article. More information about DOIs can be found at:

Open Access
If your paper is accepted, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper.

For authors signing the copyright transfer agreement
If the OnlineOpen option is not selected the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs below:

CTA Terms and Conditions

For authors choosing OnlineOpen
If the OnlineOpen option is selected the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):

Creative Commons Attribution License OAA
Creative Commons Attribution Non-Commercial License OAA
Creative Commons Attribution Non-Commercial -NoDerivs License OAA

To preview the terms and conditions of these open access agreements please visit the Copyright FAQs hosted on Wiley Author Services and visit

If you select the OnlineOpen option and your research is funded by The Wellcome Trust or members of the Research Councils UK (RCUK), you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with Wellcome Trust and Research Councils UK requirements. For more information on this policy and the Journal’s compliant self-archiving policy please visit:

World Wide Web Site

Additional material (such as multimedia adjuncts, large data sets, extra colour illustrations, bibliographies, or any other material for which insufficient space in Acta Zoologica is available) may be able to be published via the journal's World Wide Web site.