Journal of Anatomy

Cover image for Vol. 225 Issue 2

Edited By: Julia Clarke, Thomas Gillingwater, Anthony Graham, Stefan Milz

Impact Factor: 2.357

ISI Journal Citation Reports © Ranking: 2012: 5/21 (Anatomy & Morphology)

Online ISSN: 1469-7580

Associated Title(s): Aging Cell

Author Guidelines

Articles submitted to Journal of Anatomy are done so on the following conditions: that the work described has not been published previously (except in the form of an abstract or as part of a published lecture or academic thesis), that it is not under consideration for publication elsewhere, that its publication is approved by all authors and tacitly or explicitly by the responsible authorities where the work was carried out, and that, if accepted, it will not be published elsewhere including electronically in the same form, in English or in any other language, without the written consent of the copyright-holder.

There are no page charges, and charges are not made for colour, if, in the Editors’ opinion it is essential. In all other cases costs must be borne by the author. The Editors-in-Chief of Journal of Anatomy endorse the ARRIVE guidelines for reporting experiments (, and expect authors to refer to these guidelines before submission of a manuscript.

NIH-funded authors and Journal of Anatomy
The NIH mandates grantees to deposit their peer-reviewed author manuscripts in PubMed Central, to be made publicly available within 12 months of
publication. The NIH mandate applies to all articles based on research that has been wholly or partially funded by the NIH and that are accepted for publication on or after April 7, 2008. In order to help authors comply with the NIH mandate, for papers accepted for publication in Journal of Anatomy, Wiley Blackwell will post the accepted manuscript (incorporating all amendments made during peer review, but prior to the publisher's copy-editing and typesetting) of articles by NIH grant-holders to PubMed Central at the point of acceptance by the journal. This version will then be made publicly available in PubMed Central 12 months after publication. Following the deposit Wiley Blackwell authors will receive further communications from the NIH with respect to the submission. For further information, see here.

If authors wish to make their final published article openly accessible and without a 12 month embargo, they can choose to publish via the OnlineOpen service.
Wellcome and HHMI grantees can find out further information here.

Pre-submission English-language editing
Authors for whom English is a second language may choose to have their manuscript professionally edited before submission to improve the English. A list of independent suppliers of editing services can be found at . All services are paid for and arranged by the author, and use of one of these services does not guarantee acceptance or preference for publication.

Manuscript submission online (please see can be as Word document (.doc) or (.docx), or RTF (.rtf). If submitting to Manuscript Central using LaTeX please upload a PDF file of the manuscript for the reviewers. Note that upon acceptance we will require your TeX/LaTeX source files to edit and typeset the article. Figures for review should be uploaded separately as GIF (.gif), JPEG (.jpg), TIFF (.tif), or EPS (.eps). On acceptance, you will be required to provide HIGH RESOLUTION GRAPHICS FILES (note that GIF (.gif), JPEG (.jpg), and PNG (.png) files are not acceptable for publication).
Full upload instructions and support are available online from the submission site via the Get Help Now button. Please submit your covering letter or comments to the Editor in Chief when prompted online.

Revised manuscripts must be submitted in their final form, within 2 months of authors being notified of conditional acceptance (pending satisfactory revision). Resubmissions after this time will be considered as new.

Text should be divided into the following sections and appear in the order:
(1) title page (with short running page heading, title, authors names and affiliations);
(2) summary;
(3) keywords;
(4) introduction;
(5) main body;
(6) concluding remarks;
(7) acknowledgements;
(8) references;
(9) supplementary material;
(10) tables; and
(11) figure legends.
Authorities for the latin binomial of every organism are not used in the title or summary, and only on the first mention in the main body of the text. 
Gene names and loci should be italic, proteins should be roman.
Virus nomenclature (and acronyms) should follow the guidelines of the International Committee on the Taxonomy of Viruses (ICTV). The current report is: van Regenmortel MHV, Fauquet CM, Bishop DHL (Eds) (2001) Virus Taxonomy: Seventh Report of the International Committee on Taxonomy Viruses. San Diego: Academic Press. Authors are also advised to check the ITCV website for the latest information.
Chemical nomenclature should follow the International Union of Pure and Applied Chemistry (IUPAC) definitive rules for nomenclature.
Industrial products should be referred to by their common names (ISO Publications 1831, 2474, etc.). In the absence of a common name, use the full name or a defined abbreviation, in preference to a trade name. At first mention, trade names should be capitalised.
Statistics: Calculations and the validity of deductions made from them should be checked and validated by a statistician.
Symbols, units and abbreviations should be expressed as Système International (SI) units, as given in: Baron DN (Ed) (1994) Units and Abbreviations: a guide for medical authors and editors. London: Royal Society of Medicine Press. In exceptional circumstances, others may be used, provided they are consistent. Apply to the Editorial Office for advice.

Author Contributions
Journal of Anatomy recommends that authors submit a short description of all contributions to their manuscript. Each author’s contribution should be described in brief, to appear immediately before the references. Authorship credit might include contributions to concept/design, acquisition of data, data analysis/interpretation, drafting of the manuscript, critical revision of the manuscript and approval of the article. Contributors who do not qualify as authors can also be listed and their particular contribution described. Further information on authorship and contributorship may be found here.

References in the text should be inserted in parentheses in full for single and dual authored papers, but using the first author and et al. for multiple authored papers. Reference to personal communications, unedited and un-refereed work, and work that is unpublished should be minimal and should appear in the text only. It is the author’s responsibility to obtain permission from colleagues to include their work as a personal communication.
References in the list should follow the Harvard system. Over 6 authors will be abbreviated to 3 authors et al. Refer to a recent copy of the journal for examples.

We recommend the use of a tool such as EndNote or Reference Manager for reference management and formatting.

EndNote reference styles can be searched for here.
Reference Manager reference styles can be searched for here.

Supporting Information
Supporting Information can be a useful way for an author to include important but ancillary information with the online version of an article. Examples of Supporting Information include additional tables, data sets, figures, movie files, audio clips, 3D structures, and other related nonessential multimedia files. Supporting Information should be cited within the article text, and a descriptive legend should be included. It is published as supplied by the author, and a proof is not made available prior to publication; for these reasons, authors should provide any Supporting Information in the desired final format. For further information on recommended file types and requirements for submission, please visit:

Numerical results should be presented either as tables or figures, but not both. The journal welcomes colour figures and plates, when information would be lost if reproduced in black and white.
Tables: Tables should be typed on separate pages, as an integral part the text file. They should have a brief descriptive title and be self-explanatory. Units should appear in parentheses in the column headings, not in the body of the table. Repeated words or numerals on successive lines should be written in full. Footnotes should be minimal. When the precision of data is expressed as standard error (se) or standard errors of differences (sed) the degrees of freedom (df) should be given.
Electronic artwork: JAnat only accepts submission of electronic artwork. The journal accepts the following formats only:

(1) GRAPHS, CHARTS and MAPS as Encapsulated Postscript Format (.eps).
The best results are obtained with software applications that can output EPS format (Systat SigmaPlot; Adobe Illustrator; CorelDraw; Deneba Canvas; Macromedia Freehand); and

(2) PHOTOGRAPHS in Tagged Image File Format (.tif).
TIFF files should be supplied at a minimum resolution of 300 dpi (dots per inch) at the final size at which they are to appear in the journal. TIFF and EPS formats are the industry standard for archiving and print and online publication.
Detailed information on digital illustration standards is available at this webpage

Please note that file formats other than EPS and TIF will be automatically rejected by the online submission website.
Colour files should be in RGB format. Labelling should be in 10pt sans serif style font. Figure sections should be designated with upper case letters. Magnification bars should be given on electron and light micrographs.

Original artwork: If you are unable to submit electronic artwork, please provide two sets of all figures as high-quality glossy prints at the size that they are to appear in print. One set should be unlabelled.
Legends: Table and figure legends should be included within the text file and contain sufficient information to be understood without reference to the text. Each should begin with a short title for the figure. All symbols and abbreviations should be explained within the legend.
Cover images: Electronic artwork of high quality suitable for the cover of JAnat are welcomed. They should be sent to the Editorial Office and be accompanied by a relevant caption. It is preferred that images should be related to submitted papers. Contributors are required to grant exclusive publishing rights to the Society. It is the Publisher's policy not to publish items without signed copyright transfer agreement.

Authors must obtain permission to reproduce any copyright material, and include an acknowledgement of the source in their Article. Authors should be aware that the unreferenced use of the published and unpublished ideas, writing or illustrations of others, from whatever source (including research grant applications), or submission of a complete paper under 'new' authorship in the same or a different language, constitutes plagiarism.

Manuscripts are accepted on the understanding that they will not be published elsewhere in any form, without the prior consent of the Society. Your material will not be passed to the Publisher without receipt of:
1. covering letter documenting any revisions required by the editor;
2. signed copyright transfer agreement;
3. completed accepted artwork checklist - documenting the graphic files supplied in accordance with industry standards; and

Authors will be required to assign copyright to the Anatomical Society. If your paper is accepted, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper.

For authors signing the copyright transfer agreement
If the OnlineOpen option is not selected the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs below:
CTA Terms and Conditions

For authors choosing OnlineOpen
OnlineOpen is available to authors of articles who wish to make their article open access. With OnlineOpen the author, their funding agency, or institution pays a fee to ensure that the article is made available to non-subscribers upon publication via Wiley Online Library, as well as deposited in PubMed Central and PMC mirror sites. In addition to publication online via Wiley Online Library, authors of OnlineOpen articles are permitted to post the final, published PDF of their article on a website, institutional repository, or other free public server, immediately on publication.

If you want your article to be open access please choose the appropriate licence agreement when you log in to Wiley’s Author Services system. Click on ‘Make my article OnlineOpen’ and choose the appropriate license by clicking on ‘Sign license agreement now’ when you log in to Wiley’s Author Services system.

For the full list of terms and conditions, see

If the OnlineOpen option is selected the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):

Creative Commons Attribution License OAA
Creative Commons Attribution Non-Commercial License OAA
Creative Commons Attribution Non-Commercial -NoDerivs License OAA

To preview the terms and conditions of these open access agreements please visit the Copyright FAQs hosted on Wiley Author Services and visit

If you select the OnlineOpen option and your research is funded by The Wellcome Trust and members of the Research Councils UK (RCUK) or the Austrian Science Fund (FAF) you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with your Funder requirements. For more information on this policy and the Journal’s compliant self-archiving policy please visit:

Online production tracking is now available for your article through Wiley Blackwell Author Services.
Author Services enables authors to track their article – once it has been accepted – through the production process to publication online and in print. Authors can check the status of their articles online and choose to receive automated e-mails at key stages of production. The author will receive an e-mail with a unique link that enables them to register and have their article automatically added to the system. Please ensure that a complete e-mail address is provided when submitting the manuscript. Visit Wiley Blackwell Author Services for more details on online production tracking and for a wealth of resources including FAQs and tips on article preparation, submission and more.

Email notification that your work is in proof and ready for collection will be sent from the JAnat e-proofing website. Further instructions will be sent with the upload notification email. Acrobat Reader is required in order to read this file. This can be downloaded (free of charge) from Adobe
In your absence, please arrange for a colleague to correct on your behalf. Major alterations will be charged to the author and delay publication.

Free access to the final PDF offprint of your article will be available via author services only. Please therefore sign up for author services if you would like to access your article PDF offprint and enjoy the many other benefits the service offers.

Additional paper offprints may be ordered online. Please click on the following link fill in the necessary details and ensure that you type information in all of the required fields:

If you have queries about paper offprints please email

Please note that unless specifically requested, Wiley Blackwell Publishing will dispose of all hardcopy or electronic material submitted two months after publication. If you require the return of any material submitted, please inform the editorial office or production editor as soon as possible if you have not yet done so.

Last updated: 11 July 2014