Journal of Anatomy

Cover image for Vol. 230 Issue 2

Edited By: Julia Clarke, Thomas Gillingwater, Anthony Graham, Stefan Milz

Impact Factor: 2.154

ISI Journal Citation Reports © Ranking: 2015: 4/21 (Anatomy & Morphology)

Online ISSN: 1469-7580

Associated Title(s): Aging Cell

Author Guidelines

For the Journal's Aim and Scope Click here

Articles submitted to Journal of Anatomy are done so on the following conditions: that the work described has not been published previously (except in the form of an abstract or as part of a published lecture or academic thesis), that it is not under consideration for publication elsewhere, that its publication is approved by all authors and tacitly or explicitly by the responsible authorities where the work was carried out, and that, if accepted, it will not be published elsewhere including electronically in the same form, in English or in any other language, without the written consent of the copyright-holder.

There are no page charges, and charges are not made for colour, if, in the Editors’ opinion it is essential. In all other cases costs must be borne by the author. The Editors-in-Chief of Journal of Anatomy endorse the ARRIVE guidelines for reporting experiments (, and expect authors to refer to these guidelines before submission of a manuscript.

NIH-funded authors and Journal of Anatomy
The NIH mandates grantees to deposit their peer-reviewed author manuscripts in PubMed Central, to be made publicly available within 12 months of
publication. The NIH mandate applies to all articles based on research that has been wholly or partially funded by the NIH and that are accepted for publication on or after April 7, 2008. In order to help authors comply with the NIH mandate, for papers accepted for publication in Journal of Anatomy, Wiley Blackwell will post the accepted manuscript (incorporating all amendments made during peer review, but prior to the publisher's copy-editing and typesetting) of articles by NIH grant-holders to PubMed Central at the point of acceptance by the journal. This version will then be made publicly available in PubMed Central 12 months after publication. Following the deposit Wiley Blackwell authors will receive further communications from the NIH with respect to the submission. For further information, see here.

If authors wish to make their final published article openly accessible and without a 12 month embargo, they can choose to publish via the OnlineOpen service.
Wellcome and HHMI grantees can find out further information here.

Pre-submission English-language editing
Authors for whom English is a second language may choose to have their manuscript professionally edited before submission to improve the English. A list of independent suppliers of editing services can be found at . All services are paid for and arranged by the author, and use of one of these services does not guarantee acceptance or preference for publication.

Types of manuscripts
The majority of manuscripts published in Journal of Anatomy are Original Articles. There are no limits regarding the word count or number of display figures for Original Articles, but authors are encouraged to write in a concise manner. Authors are discouraged from submitting papers longer than 70 manuscript pages (including figures and tables). The Editors will also consider shorter manuscripts for publication as Brief Communications. These papers are normally shorter than 15 manuscript pages and contain no more than three display figures. In addition, the Editors will consider manuscripts reporting on methodological developments, for publication as Methods papers (either as full length manuscripts or as Brief Communications).

Review articles are normally commissioned by the Editors, but we welcome pre-submission enquiries (prospective authors are encouraged to email a copy of the proposed title and abstract to Similarly, Historical Perspective and Educational Perspective manuscripts are normally commissioned by the Editors, but again we welcome pre-submission enquiries (prospective authors are encouraged to email a copy of the proposed title and abstract to

For all manuscripts, during the online submission process, authors will be asked to choose a subject area (these are only used to assist with the allocation of appropriate editors and reviewers) that best represents their paper, from the following:

Cell biology and tissue architecture
Comparative functional morphology
Developmental biology
Evolutionary developmental biology
Evolutionary morphology
Functional human anatomy
Integrative vertebrate paleontology
Methodological innovations in anatomical research
Musculoskeletal system
Neuroanatomy and neurodegeneration
Significant advances in anatomy education

There is an additional category, ‘Symposium papers’, but this should only be used where an author has been specifically invited by one of the Editors in Chief or by a Guest Editor to submit a paper to a forthcoming special symposium issue.

Online manuscript submission
Submission of a manuscript to Journal of Anatomy implies that it reports unpublished work and that it is not under consideration for publication elsewhere. If previously published tables, illustrations or more than 200 words of text are to be included in the manuscript, then the copyright holder's permission must be obtained. Copies of permission letters should be included with the manuscript.

Manuscripts should be submitted online at Full upload instructions and support are available online from the submission site via the Get Help Now button.

Manuscripts can be uploaded as a Word document (.doc or .docx), or as a Rich Text Format document (.rtf). If submitting to Manuscript Central using LaTeX please upload a PDF file of the manuscript for the reviewers. Note that upon acceptance we will require your TeX/LaTeX source files to edit and typeset the article. Figures for review should be uploaded separately as GIF (.gif), JPEG (.jpg), TIFF (.tif), or EPS (.eps). On acceptance, you will be required to provide HIGH RESOLUTION GRAPHICS FILES (note that GIF (.gif), JPEG (.jpg), and PNG (.png) files are not acceptable for publication).

Please submit a covering letter addressed to the Editors when prompted online. The covering letter should state the importance and anatomical relevance of the study. Authors will also be asked to provide the names of one or more preferred referees (who have no personal or professional connection with the study or the authors), although the Editors are not obliged to use these.

Review of manuscripts

Due to space restrictions and to the volume of high quality papers submitted, the Editors reserve the right to return immediately those papers that are unlikely to be competitive for space in the journal and/or those that do not conform to the general editorial philosophy and standards of Journal of Anatomy. Wherever possible, such editorial decisions will be processed within a week of submission. Manuscripts selected for external peer review will normally be reviewed within four weeks of submission.

Authors should refer to recently published manuscripts for an overall guide to Journal style (available at

Text should be divided into the following sections and appear in the order:
(1) title page (with short running page heading, title, authors names and affiliations);
(2) abstract (no longer than 500 words, in a single paragraph, not divided into sections with headings);
(3) keywords;
(4) main body: introduction, methods, results, discussion (this may be combined with the results section) and conclusions if desired;
(5) acknowledgements;
(6) author contributions (see below);
(7) references;
(8) supplementary material (if any);
(9) tables; and
(10) figure legends.
Please avoid the use of footnotes, and number the pages using the page numbering feature of the word processing software.

Authorities for the Latin binomial of every organism are not used in the title or summary, and only on the first mention in the main body of the text.
Gene names and loci should be in italic text and protein names should use in roman text.

Virus nomenclature (and acronyms) should follow the guidelines of the International Committee on the Taxonomy of Viruses (ICTV). The current report is: van Regenmortel MHV, Fauquet CM, Bishop DHL (Eds) (2001) Virus Taxonomy: Seventh Report of the International Committee on Taxonomy Viruses. San Diego: Academic Press. Authors are also advised to check the ITCV website for the latest information.

Chemical nomenclature should follow the International Union of Pure and Applied Chemistry (IUPAC) definitive rules for nomenclature.

Industrial products should be referred to by their common names (ISO Publications 1831, 2474, etc.). In the absence of a common name, use the full name or a defined abbreviation, in preference to a trade name. At first mention, trade names should be capitalised.

Statistics: Calculations and the validity of deductions made from them should be checked and validated by a statistician.

Symbols, units and abbreviations should be expressed as Système International (SI) units, as given in: Baron DN (Ed) (1994) Units and Abbreviations: a guide for medical authors and editors. London: Royal Society of Medicine Press. In exceptional circumstances, others may be used, provided they are consistent. Apply to the Editorial Office for advice.

Author contributions

Journal of Anatomy recommends that authors submit a short description of all contributions to their manuscript. Each author’s contribution should be described in brief, to appear immediately before the references. Authorship credit might include contributions to concept/design, acquisition of data, data analysis/interpretation, drafting of the manuscript, critical revision of the manuscript and approval of the article. Contributors who do not qualify as authors can also be listed and their particular contribution described. Further information on authorship and contributorship may be found here.


References in the text should be inserted in parentheses in full for single and dual authored papers, but using the first author and et al. for multiple authored papers. Reference to personal communications, unedited and un-refereed work, and work that is unpublished should be minimal and should appear in the text only. It is the author’s responsibility to obtain permission from colleagues to include their work as a personal communication.
References in the list should follow the Harvard system. Over 6 authors will be abbreviated to 3 authors et al. Refer to a recent copy of the journal for examples.

We recommend the use of a tool such as EndNote or Reference Manager for reference management and formatting.

Supporting Information

Supporting Information can be a useful way for an author to include important but ancillary information with the online version of an article. Examples of Supporting Information include additional tables, data sets, figures, movie files, audio clips, 3D structures, and other related nonessential multimedia files. Supporting Information should be cited within the article text, and a descriptive legend should be included. It is published as supplied by the author, and a proof is not made available prior to publication; for these reasons, authors should provide any Supporting Information in the desired final format. For further information on recommended file types and requirements for submission, please visit:

Numerical results should be presented either as tables or figures, but not both. The Journal welcomes colour figures and plates, when information would be lost if reproduced in black and white.

Tables: Tables should be typed on separate pages, as an integral part the text file. They should have a brief descriptive title and be self-explanatory. Units should appear in parentheses in the column headings, not in the body of the table. Repeated words or numerals on successive lines should be written in full. Footnotes should be minimal. When the precision of data is expressed as standard error (se) or standard errors of differences (sed) the degrees of freedom (df) should be given.

Electronic artwork: Journal of Anatomy only accepts submission of electronic artwork. The journal accepts the following formats only:

(1) GRAPHS, CHARTS and MAPS as Encapsulated Postscript Format (.eps).
The best results are obtained with software applications that can output EPS format (Systat SigmaPlot; Adobe Illustrator; CorelDraw; Deneba Canvas; Macromedia Freehand); and

(2) PHOTOGRAPHS in Tagged Image File Format (.tif).
TIFF files should be supplied at a minimum resolution of 300 dpi (dots per inch) at the final size at which they are to appear in the journal. TIFF and EPS formats are the industry standard for archiving and print and online publication.

Detailed information on digital illustration standards is available at this webpage.

Please note that file formats other than EPS and TIF will be automatically rejected by the online submission website.

Colour files should be in RGB format. Labelling should be in 10pt sans serif style font. Figure sections should be designated with upper case letters. Magnification bars should be given on electron and light micrographs.

Legends: Table and figure legends should be included within the text file and contain sufficient information to be understood without reference to the text. Each should begin with a short title for the figure. All symbols and abbreviations should be explained within the legend.

Cover images: Electronic artwork of high quality suitable for the cover of Journal of Anatomy are welcomed. They should be sent to the Editorial Office ( and be accompanied by a relevant caption. It is preferred that images should be related to submitted papers. Contributors are required to grant exclusive publishing rights to the Society. It is the Publisher's policy not to publish items without signed copyright transfer agreement.


Authors must obtain permission to reproduce any copyright material, and include an acknowledgement of the source in their Article. Authors should be aware that the unreferenced use of the published and unpublished ideas, writing or illustrations of others, from whatever source (including research grant applications), or submission of a complete paper under 'new' authorship in the same or a different language, constitutes plagiarism.


Manuscripts are accepted on the understanding that they will not be published elsewhere in any form, without the prior consent of the Society. Your material will not be passed to the Publisher without receipt of:
1. covering letter documenting any revisions required by the editor;
2. signed copyright transfer agreement;
3. completed accepted artwork checklist - documenting the graphic files supplied in accordance with industry standards; and
5. Any potential or perceived conflict of interest must be declared in the Acknowledgements section of the manuscript. If no conflict exists, please add a sentence to this effect in the Acknowledgements section.


Authors will be required to assign copyright to the Anatomical Society. If your paper is accepted, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper.

For authors signing the copyright transfer agreement

If the OnlineOpen option is not selected the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs below:
CTA Terms and Conditions

For authors choosing OnlineOpen
OnlineOpen is available to authors of articles who wish to make their article open access. With OnlineOpen the author, their funding agency, or institution pays a fee to ensure that the article is made available to non-subscribers upon publication via Wiley Online Library, as well as deposited in PubMed Central and PMC mirror sites. In addition to publication online via Wiley Online Library, authors of OnlineOpen articles are permitted to post the final, published PDF of their article on a website, institutional repository, or other free public server, immediately on publication.

If you want your article to be open access please choose the appropriate licence agreement when you log in to Wiley’s Author Services system. Click on ‘Make my article OnlineOpen’ and choose the appropriate license by clicking on ‘Sign license agreement now’ when you log in to Wiley’s Author Services system.

For the full list of terms and conditions, see

If the OnlineOpen option is selected the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):

Creative Commons Attribution License OAA
Creative Commons Attribution Non-Commercial License OAA
Creative Commons Attribution Non-Commercial -NoDerivs License OAA

To preview the terms and conditions of these open access agreements please visit the Copyright FAQs hosted on Wiley Author Services and visit

If you select the OnlineOpen option and your research is funded by The Wellcome Trust and members of the Research Councils UK (RCUK) or the Austrian Science Fund (FAF) you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with your Funder requirements. For more information on this policy and the Journal’s compliant self-archiving policy please visit:


Online production tracking is now available for your article through Wiley Blackwell Author Services.
Author Services enables authors to track their article – once it has been accepted – through the production process to publication online and in print. Authors can check the status of their articles online and choose to receive automated e-mails at key stages of production. The author will receive an e-mail with a unique link that enables them to register and have their article automatically added to the system. Please ensure that a complete e-mail address is provided when submitting the manuscript. Visit Wiley Blackwell Author Services for more details on online production tracking and for a wealth of resources including FAQs and tips on article preparation, submission and more.


Email notification that your work is in proof and ready for collection will be sent from the Journal of Anatomy e-proofing website. Further instructions will be sent with the upload notification email. Acrobat Reader is required in order to read this file. This can be downloaded (free of charge) from Adobe.
In your absence, please arrange for a colleague to correct on your behalf. Major alterations will be charged to the author and delay publication.


Free access to the final PDF offprint of your article will be available via author services only. Please therefore sign up for author services if you would like to access your article PDF offprint and enjoy the many other benefits the service offers.
Additional paper offprints may be ordered online. Please click on the following link fill in the necessary details and ensure that you type information in all of the required fields:
If you have queries about paper offprints please email
Please note that unless specifically requested, Wiley Blackwell Publishing will dispose of all hardcopy or electronic material submitted two months after publication. If you require the return of any material submitted, please inform the editorial office or production editor as soon as possible if you have not yet done so.

Last updated: 30 September 2015