Entomological Science

Cover image for Vol. 19 Issue 4

Edited By: Masanori J. Toda

Impact Factor: 1.144

ISI Journal Citation Reports © Ranking: 2015: 39/94 (Entomology)

Online ISSN: 1479-8298

Author Guidelines

Authors wishing to submit to Entomological Science should read these instructions carefully before preparing their manuscripts. They are also recommended to consult the following reference: Scientific Style and Format: The CBE Manual for Authors, Editors, and Publishers, Eighth Edition, CBE Style Manual Committee, 1994, Cambridge University Press (http://www.scientificstyleandformat.org/Home.html).


Entomological Science is the official English language journal of the Entomological Society of Japan. The Journal publishes original research papers and reviews from any entomological discipline or from directly allied field in ecology, behavioral biology, physiology, biochemistry, development, genetics, systematics, morphology, evolution and general entomology. Papers of applied entomology will be considered for publication if they significantly advance the field of entomological science in the opinion of the Editors and Editorial Board.

Authors should submit their manuscripts to Entomological Science online to facilitate even quicker and more efficient processing. Please log onto the sitedirectly at: http://mc.manuscriptcentral.com/ens

All manuscripts submitted to the Journal must comply with these Author Guidelines. Failure to do so will result in return of the manuscript and possible delay in publication.

Manuscripts should be written so that they are intelligible to the professional reader who is not a specialist in the particular field. They should be written in English, in a clear, concise, direct style. Where contributions are judged as acceptable for publication on the basis of scientific content, the Editor and the Publisher reserve the right to modify typescripts to eliminate ambiguity and repetition and improve communication between author and reader.

Pre-submission English-language editing
Authors for whom English is a second language may choose to have their manuscript professionally edited before submission. A list of independent suppliers of editing services can be found at
http://authorservices.wiley.com/bauthor/english_language.asp. All services should be paid for and arranged by the author, and the use of these services does not guarantee acceptance or preference for publication.

Manuscripts may be submitted as research papers or review articles. Research papers should be the product of original scientific research or observation, and are classed as either Original Articles or Short Communications. Short Communications should have the main body of the text (excepting title, authors’ information, abstract, key words, acknowledgments and reference list) no longer than three printed pages (one page approximately equivalent to 770 words or 5100 letters including spaces of solid text).

All manuscripts, including short communications, should include a discussion of the significance of the results and their relationship to other work, and will be reviewed by at least two referees. Manuscripts will be considered for publication only if they have not been published or submitted for publication elsewhere.


File formats and page settings
Manuscripts must be on A4 size pages, in 12 pt font size, with 24 to 28 lines per page. The manuscript has to be submitted as Microsoft Word or Rich Text Format.

A submission template in Microsoft Word is available for Original Article and Short Communication.

Manuscript composition
The manuscript should start with the title of the paper, followed by the names of the authors, the addresses of the institutions at which the work was carried out, an abstract, key words and the main body of text. The abstract should be a concise summary (250 words or fewer) of the manuscript's significant content. There should be three to seven key words, which should not duplicate words in the title and should be arranged in alphabetical order, and preferably include the order and family names of the major organism(s) considered, if not appearing in the title.

The IMRAD format ("Introduction", "Materials and methods", "Results" and "Discussion") is recommended for Original Articles, although following this format is at the author's discretion. The main body of text in Short Communications should not be divided into sections with headings.

Character and paragraph formats
Words to be italicized should be typed in italics but should not be underlined, and capitalization should be avoided in all cases except for headings, abbreviations of depositories and/or institutions, and technical terms. Hard returns should be used only at the end of a paragraph and words should not be hyphenated. Other formatting, such as headings and subheadings, should follow that in the latest issue of Entomological Science.

Units and numeric values
Measurements should be expressed in units of SI (International System of Units). A space should be inserted between number and unit except ˚C and %., and before and after =, ±, and so on. Numbers between one and ten in the main body of the text should be written in full except dates, the numbering of figures, numbers accompanying metrical units (such as mm, m, km, mg, g, kg, s, min and h), and numbers used in taxonomic descriptions. Numbers greater than ten should be written as numerals. Numbers greater than 1000 should not include any thousand separators. Fractions should be written as decimals.

Scientific names
All papers must conform to the latest edition of the International Code of Zoological Nomenclature. The first mention of an animal should include the full scientific name with the authority and, particularly in taxonomic papers, the year of publication. The authority and the year of publication should be separated by a comma. Genus names should not be abbreviated at the beginning of a sentence.

Research permits
If the research was carried out in areas for which research permits are necessary (e.g. nature reserves) or if it deals with organisms for which collection or import/export permits are required, the authors must clearly detail obtaining these permits in the acknowledgments paragraph.

Taxonomic papers
In taxonomic papers, type specimens must be clearly designated and type depositories must be clearly indicated. Authors are required to deposit the name-bearing type material in an internationally recognized institution (not in private collections). For the number of segment, Roman numerals (e.g. tergum X) should be used. Body part terms specific to a given taxon should be explained in the "MATERIALS AND METHODS" or be indicated in figures. The second couplets of the key should start with a dash only (‒). The list of specimens examined should be as compact as possible; for example, "Specimens examined. Japan: 1♀ (IUNH), Mito, 26.xi.2004, J. Kojima; 3♂2♀ (IUNH), . . ." The holotypes (or other name-bearing types) and paratypes (or paralectotypes) are listed separately, and full label data should be given for the name-bearing types in quotation marks when they are designated in the paper. All publications including new nomenclatural acts will be registered to ZooBank by the Editorial Office before online publication.

Acknowledgments should be typed in a single paragraph headed "ACKNOWLEDGMENTS", directly preceding the "REFERENCES" section. The source of financial grants and other funding must be acknowledged, including a frank declaration of the authors' industrial links and affiliations. The contribution of colleagues or institutions should also be acknowledged. Personal thanks and thanks to editors and anonymous reviewers are not appropriate.

The Harvard (author/date) system of referencing should be used. In the text, give the author's name followed by the year in parentheses: Smith (2000). If there are two authors, use 'and': Smith and Jones (2001); but if cited within parentheses use '&': (Smith & Jones 2001). When reference is made to a work by three or more authors, the first name followed by et al. should be used: MacDonald et al. (2002). Multiple references within the same parentheses should be chronologically ordered and be separated with semicolon. In taxonomic papers references in synonymic lists should be cited in the following style: Drosophila melanogaster Meigen, 1830: 85; Kikkawa & Peng, 1938: 534.

In the reference list, references should be listed in alphabetical order of the first authors. The order of references of the same first author is as follows: (1) single-authored references should be chronologically ordered; (2) references of two authors should be listed in alphabetical order of the second author; (3) references of three or more authors should be chronologically ordered (those of the same year should be discriminated by adding a letter of ‘a’, ‘b’, …, even though co-authors are different). Cite the names of all authors (or editors) when there are six or fewer; when seven or more, list the first three followed by et al. Do not use ibid. or op cit. The title of the cited work should be given in full and journal titles should not be abbreviated. Reference to unpublished data and personal communications should be avoided if at all possible, but if absolutely necessary should be cited in the text only, e.g. (A. Smith, unpubl. data, 2000). All other citations in the text (including synonymic lists), tables and figures must be listed in the reference list.

Journal articles with sequential page numbers of volume
Atkinson WD, Shorrocks B (1984) Aggregation of larval Diptera ... for coexistence. American Naturalist 124, 336‒351. (Use a ‘dash’ but not ‘hyphen’ as the connector of page numbers.)

Journal articles without sequential page numbers of volume
Sugime Y, Ogawa K, Watanabe D, Shimoji H, Koshikawa S, Miura T (2015) Expansion of presoldier cuticle contributes to head elongation during soldier differentiation in termites. The Science of Nature 102, Article ID 71. DOI: 10.1007/s00114-015-1322-3

Journal articles published online in early view
Kohyama TI, Matsubayashi KW, Katakura H (2015) Heterospecific sperm reduction in interspecific crosses between two closely related phytophagous ladybird beetles, Henosepilachna vigintioctomaculata and H. pustulosa (Coleoptera: Coccinellidae). Entomological Science. Published online: 4 Dec 2015; DOI: 10.1111/ens.12159

Gerson U, Simley R (1990) Acarine Biocontrol Agents: An Illustrated Key and Manual. Chapman and Hall, London.

Chapter in a book
Weis AE (1992) Plant variation ... herbivore performance. In: Fritz RS, Simms EL (eds) Plant ... and Pathogens, pp 140‒171. University of Chicago Press, Chicago, IL.

Title in English but text in a language other than English
Matsuura M (1995) Social Wasps of Japan in Color. Hokkaido University Press, Sapporo. (In Japanese.)

Title translated by the author(s) into English
Tanaka A (1990) [Feeding Habits of Diptera]. Hokuryukan, Tokyo. (In Japanese.)

Sorenson MD, Franzosa EA (2007) TreeRot, Version 3. Boston University, Boston, MA. Available from URL: http://people.bu.edu/msoren/TreeRot.html

Japanese Ant Database Group (2003) Japanese Ant Image Database 2003 [updated 19 Oct 2006; cited 15 Dec 2015]. Available from URL: http://ant.edb.miyakyo-u.ac.jp/E/index.html

We recommend the use of a tool such as Reference Manager for reference management and formatting. Reference Manager reference styles can be searched for here: http://www.refman.com/support/rmstyles.asp


Tables and figures should be limited to those required for clear communication and must be referred to: in the text, Table 1, Tables 1 and 2, Tables 1‒3, Figure 1, Figures 1 and 2, Figures 1‒3; within parentheses, (Table 1), (Tables 1,2), (Tables 1‒3), (Fig. 1), (Fig. 1A), (Fig. 1A,B), (Figs 1,2), (Figs 1A,2A), (Figs 1‒3). The same data should not be presented in both table and figure formats. Each table should be typed with title (one sentence) and footnotes (if necessary) on a separate page (or multiple pages if required) at the end of main text or as a separate file, and be numbered in sequence with Arabic characters. In electronic files, tables should be prepared in editable formats of MS Word or Excel; pasted images are not acceptable.

All illustrations, including photographs, graphs and maps are treated as figures, even if they are arranged in plates. Figures should be arranged in size no larger than 170 x 220 mm and, if possible, they should be sized to fit within a single column (up to 80 mm) or full text width (up to 170 mm) in order to utilize the paper layout. Publication of color figures is free of charge. Authors may be asked to restructure the figures in plate by order of Editors if some are considered to be smaller.

All lettering in the figures, where possible, should be in 'Arial' font and 12 pt or bigger.

The degree of magnification used should be indicated by means of a scale line.

Figure legends should be included with the main text of the manuscript, after the references on separate page(s).

For digital figure files format, 300 dpi TIFF in halftone images and EPS or 800 dpi TIFF in line arts or combination figures are recommended. If the provided files are too low in resolution or are problematic, authors may be asked for a resupply after acceptance.

Wiley websites for authors also provide detailed information on the preparation and submission of articles at http://authorservices.wiley.com/bauthor/journal.asp, and figures at http://authorservices.wiley.com/bauthor/illustration.asp

Supporting Information
Authors are welcome to submit additional ”Supporting Information”, such as data sets or additional figures or tables. This material will not be published in the print edition of the journal but it will be viewable via the online edition. Only material that is a valuable addition to the article should be included. Such supporting information should be referred to in the text as, for example,”see Table S1”.
Supporting files are hosted by the Publisher in the format supplied by the author and are not copy-edited by the Publisher. It is the responsibility of the author to supply supporting information in an appropriate file format and to ensure that it is accurate and correct. Prior to publication, authors will be sent the URL of their Supporting Information for them to check the content. Extensive editing of material is not possible at this stage and the author has the responsibility to ensure that material is sent in a correct form at the time of submission.

Authors should include a ”Supporting Information” section immediately after their References section, which should be in the following form:

Additional Supporting Information may be found in the online version of this article at the publisher’s web-site:

Appendix S1 Characters used for the parsimony analysis.
Table S1 A list of species studied.
Figure S1 Holotype and allotype of Drosophila (Siphlodora) pengi.

Please note: Wiley is not responsible for the content or functionality of any supplementary materials supplied by the authors. Any queries (other than missing material) should be directed to the corresponding author for the article.

For information on how to submit files, please click Here.

Entomological Science is committed to integrity in scientific research and recognizes the importance of maintaining the highest ethical standards. Please find the following Best Practice Guidelines on Publication Ethics. Further information at

English: http://authorservices.wiley.com/bauthor/publicationethics.asp

All submitted manuscripts are screened for possible plagiarism with the detection tool CrossCheck, which compares submitted manuscript against a database of published papers (as well as web pages) and reports the extent of similarity to previously published works.

It is Journal policy that a page charge fee is levied on articles appearing in the Journal that are in excess of 16 printed pages for Original Articles and unsolicited Review Articles. No page charge is applied to Invited Review Articles.

MEMBERS (The Entomological Society of Japan) at least one author of the article: For manuscripts exceeding this limit a page charge of ¥8 000/US$65 per each additional page is levied.

NON-MEMBERS:For manuscripts exceeding this limit a page charge of ¥12 000/US$100 per each additional page is levied. This procedure notwithstanding, no paper will be rejected or given extraordinary treatment on any basis other than its scientific merit. Any articles received by Wiley with page charges will not be published until the form has been returned.

Membership in The Entomological Society of Japan is open to anyone who has interest in entomology. Please visit the society's website at: http://www.entsoc.jp


Accepted papers will be passed to Wiley’s production team for publication. The author identified as the formal corresponding author for the paper will receive an email prompting them to login into Wiley’s Author Services, where via the Wiley Author Licensing Service (WALS) they will be asked to complete an electronic license agreement on behalf of all authors on the paper.

Authors may choose to publish under the terms of the journal’s standard copyright transfer agreement (CTA), or under open access terms made available via Wiley OnlineOpen.

Standard Copyright Transfer Agreement: FAQs about the terms and conditions of the standard CTA in place for the journal, including standard terms regarding archiving of the accepted version of the paper, are available at: Copyright Terms and Conditions FAQs.

Note that in signing the journal’s licence agreement authors agree that consent to reproduce figures from another source has been obtained.

OnlineOpen – Wiley’s Open Access Option: OnlineOpen is available to authors of articles who wish to make their article freely available to all on Wiley Online Library under a Creative Commons license. With OnlineOpen, the author, the author's funding agency, or the author's institution pays a fee to ensure that the article is made open access. Authors of OnlineOpen articles are permitted to post the final, published PDF of their article on their personal website, and in an institutional repository or other free public server immediately after publication. All OnlineOpen articles are treated in the same way as any other article. They go through the journal's standard peer-review process and will be accepted or rejected based on their own merit.

ESJ member discount on OnlineOpen fee is available from 2016 when one of authors of an article in Entomological Science is the society member. For further information and the discount code, please contact the editorial office at ent-sci@agr.hokudai.ac.jp

OnlineOpen licenses. Authors choosing OnlineOpen retain copyright in their article and have a choice of publishing under the following Creative Commons License terms: Creative Commons Attribution License (CC BY); Creative Commons Attribution Non-Commercial License (CC BY NC); Creative Commons Attribution Non-Commercial-NoDerivs License (CC BY NC ND). To preview the terms and conditions of these open access agreements please visit the Copyright Terms and Conditions FAQs.

Funder Open Access and Self-Archiving Compliance: Please click here for more information on Wiley’s compliance with specific Funder Open Access and Self Archiving Policies, and click here for more detailed information specifically about Self-Archiving definitions and policies.


Wiley’s Author Services

Author Services enables authors to track their article through the production process to publication online and in print. Authors can check the status of their articles online and choose to receive automated e-mails at key stages of production. The corresponding author will receive a unique link that enables them to register and have their article automatically added to the system. Please ensure that a complete e-mail address is provided when submitting the manuscript. Visit Author Services (https://authorservices.wiley.com/bauthor/default.asp) for more details on online production tracking and for a wealth of resources including FAQs and tips on article preparation, submission and more.


Once the paper has been typeset the corresponding author will receive an e-mail alert containing instructions on how to provide proof corrections to the article. It is therefore essential that a working e-mail address is provided for the corresponding author. Proofs should be corrected carefully; responsibility for detecting errors lies with the author.

Early View

The journal offers rapid speed to publication via Wiley’s Early View service. Early View articles are complete full-text articles published online in advance of their publication in a printed issue. Early View articles are complete and final. They have been fully reviewed, revised and edited for publication, and the authors' final corrections have been incorporated. Because they are in final form, no changes can be made after online publication. Early View articles are given a Digital Object Identifier (DOI), which allows the article to be cited and tracked before allocation to an issue. After print publication, the DOI remains valid and can continue to be used to cite and access the article. More information about DOIs can be found at http://www.doi.org/faq.html.


Article PDF for authors

A PDF of the article will be made available to the corresponding author via Author Services.

Printed Offprints

Printed offprints may be ordered online for a fee. Please click on the following link and fill in the necessary details and ensure that you type information in all of the required fields: http://offprint.cosprinters.com/cos. If you have queries about offprints please e-mail: offprint@cosprinters.com.


Editorial Office of Entomological Science,
Hokkaido University Museum,
Hokkaido University, Kita Ku, Sapporo 060-0810
Tel: +81-11-706-3739
Fax: +81-11-706-4029
Email: ent-sci@agr.hokudai.ac.jp