Invertebrate Biology

Cover image for Vol. 136 Issue 2

Edited By: Michael W. Hart

Impact Factor: 0.949

ISI Journal Citation Reports © Ranking: 2016: 76/105 (Marine & Freshwater Biology); 90/162 (Zoology)

Online ISSN: 1744-7410

Author Guidelines

Invertebrate Biology (IB) invites papers describing original, significant research focused on understanding any aspect of the biology of invertebrate organisms (protozoan and metazoan), including morphology and ultrastructure; genetics, phylogenetics, and evolution; physiology and ecology; neurobiology, behavior, and biomechanics; reproduction and development; and cell and molecular biology. Taxonomic descriptions may be a component, though secondary, of papers submitted to IB. Review articles are welcomed, but please consult the Editor-in-Chief before submitting. All contributions undergo a thorough process of peer-review.

Effective with the 2017 volume, this journal will be published in an online-only format.
Print subscription and single issue sales are available from Wiley’s Print-on-Demand Partner. To order online click through to the ordering portal from the journal’s subscribe and renew page on WOL.

Manuscript submission and format
Manuscripts should be submitted electronically through ScholarOne Manuscripts at Manuscripts must be in English with metric units throughout. For matters of style not covered here, refer to recent issues of IB or the CBE Manual Scientific Style and Format. Manuscript text files should be double-spaced, with margins of at least 2.5 cm. Please provide full names and addresses for each author, five additional keywords not in the title, and a running title of no more than 40 characters. The abstract, summarizing the main findings, conclusions, and significance of work, should be concise and informative; avoid vague statements such as "The significance of the results is discussed." The introduction, usually a brief account of background and goals, is not titled. Subsequent sections bear titles, usually Methods, Results, Discussion, and References. Subheadings should be no more than 40 characters. Place Acknowledgments between Discussion and References. Figure legends, tables, and footnotes (in that order) should follow the last page of references.

OnlineOpen is available to authors of primary research articles who wish to make their article available to non-subscribers on publication, or whose funding agency requires grantees to archive the final version of their article. With OnlineOpen, the author, the author's funding agency, or the author's institution pays a fee to ensure that the article is made available to non-subscribers upon publication via Wiley Online Library, as well as deposited in the funding agency's preferred archive.
Prior to acceptance, there is no requirement to inform an Editorial Office that you intend to publish your paper OnlineOpen if you do not wish to. All OnlineOpen articles are treated in the same way as any other article. They go through the journal's standard peer-review process and will be accepted or rejected based on their own merit.

If your paper is accepted, the author identified as the formal corresponding author for the paper will receive an email prompting them to login into Author Services; where via the Wiley Author Licensing Service (WALS) they will be able to complete the license agreement on behalf of all authors on the paper.

For authors signing the copyright transfer agreement
If the OnlineOpen option is not selected, the corresponding author will be presented with the copyright transfer agreement (CTA) to sign. The terms and conditions of the CTA can be previewed in the samples associated with the Copyright FAQs at

For authors choosing OnlineOpen
If the OnlineOpen option is selected, the corresponding author will have a choice of the following Creative Commons License Open Access Agreements (OAA):
Creative Commons Attribution License OAA
Creative Commons Attribution Non-Commercial License OAA
Creative Commons Attribution Non-Commercial -NoDerivs License OAA
To preview the terms and conditions of these open access agreements please visit the Copyright FAQs hosted on Wiley Author Services and visit If you select the OnlineOpen option and your research is funded by The Wellcome Trust and members of the Research Councils UK (RCUK), you will be given the opportunity to publish your article under a CC-BY license supporting you in complying with Wellcome Trust and Research Councils UK requirements. For more information on this policy and the Journal’s compliant self-archiving policy please visit:

Citations and references
Complete data for all published works and theses cited, and only those cited, must be listed in References, in alphabetical order; include papers accepted for publication (Baker, in press), but not those merely submitted or in preparation. In the text, cite works in chronological order: (Smith & Jones 1991; Cook et al. 1992; Anderson 1993a,b). Cite unpublished data and manuscripts from one of the authors (Cox, unpubl. data) or other individuals (D. E. Fox, pers. comm.) with no entry in References. Consult BIOSIS for journal-title abbreviations.

Examples of References style:
Grell KG & Ruthmann A 1991. Placozoa. In: Microscopic Anatomy of Invertebrates, Vol.2. Placozoa, Porifera, Cnidaria, and Ctenophora. Harrison FW & Westfall JA, eds., pp. 13–27. Wiley-Liss, New York. Klauser MD 1982. An ultrastructural and experimental study of locomotion in Trichoplax adhaerens (Placozoa). Masters thesis, Clemson University, Clemson, South Carolina. 49 pp.
Lowenstam HA & Weiner S 1989. On Biomineralization. Oxford University Press, New York. 324 pp.
Pearse VB 1989a. Growth and behavior of Trichoplax adhaerens: first record of the phylum Placozoa in Hawaii. Pac. Sci. 43: 117–121.
---- 1989b. Stalking the wild placozoan: biogeography and ecology of Trichoplax in the Pacific. (Abstr.) Am. Zool. 29: 175A.

Whenever practical, data should be presented in graphs rather than tables. In designing tables, figures, and multiple-figure plates, keep in mind that the final page size (print area) is 41 picas wide (nearly 175 mm or 7 inches) and 54 picas high (230 mm or 9 inches). Figures may occupy from half- to full-page width. Details of all figures (graphs, line drawings, halftones) must be large enough (at least 1.5 mm high), to remain clear at final, published size. Include a scale bar in each figure and its value in the figure or the legend; do not use expressions of magnification. Illustrations should be submitted as electronic files at the appropriate resolution and the size intended for publication. Color images should be submitted in CMYK color. For file format, submit line artwork (vector graphics) as Encapsulated PostScript (.eps), and halftones or photographic images (bitmap files) as Tagged Image File Format (.tif). Minimum acceptable resolution at final, published size is 300 dpi for color figures, 500 dpi for grayscale images, and 1200 dpi for line art. If a plate contains more than one type of art (e.g., line art and grayscale photographic images), the entire plate must be submitted at the highest applicable resolution (in this case, 1200 dpi). For uploading large files more quickly, LZW compression will reduce file size without affecting quality. Further information on preparing electronic figures is available at Figures and tables must be cited in the manuscript text in the same order as their numbering (e.g., Fig. 1 should be cited before Fig. 2; likewise, Fig. 1A should be cited before Fig. 1B).Authors are encouraged to submit additional, unlabeled photographs or drawings to be considered for the issue cover.

Scientific names
For all species studied, the complete scientific name with taxonomic author and date (e.g., Ostrea edulis LINNAEUS 1758) should be given either at the first mention in the text of the paper or in the Methods, but not in the title or abstract. Place taxonomic author in small caps, to distinguish from citations to reference works. Use the full binomial (Ostrea edulis) at the first mention in each section of the paper, and then abbreviate (O. edulis, not Ostrea unless referring to the genus). The Latin name of any taxon is capitalized and treated as a singular noun, not a plural or an adjective.

Instructions for authors are also available online at: and We encourage authors to support the society that publishes Invertebrate Biology, and receive the journal in which their manuscript appears, by becoming a member of the American Microscopical Society ( or