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On aims and methods of Ethology

N. Tinbergen

Department of Zoology, University of Oxford

Dedicated to Professor Konrad Lorenz at the occasion of his 60th birthday.Search for more papers by this author
First published: January‐December 1963
Cited by: 1397

Zusammenfassung

Ich habe in diesem Aufsatz kurz anzudeuten versucht, was meiner Ansicht nach das Wesentliche in Fragestellung und Methode der Ethologie ist und weshalb wir in Konrad Lorenz den Begründer moderner Ethologie erblicken. Hierbei habe ich vielleicht das Arbeitsgebiet der Ethologie weiter gefaßt, als unter Ethologen gebräuchlich ist. Wenn man aber die vielartige Arbeit jener Forscher übersieht, die sich Ethologen nennen, ist man zu dieser weiten Fassung geradezu gezwungen. Ich habe in meiner Darstellung weder Vollständigkeit noch Gleichgewicht angestrebt und, um zur Fortführung des Gesprächs anzuregen, ruhig meine Steckenpferde geritten, vor allem das Verhältnis zwischen Ethologie und Physiologie, die Gefahr der Vernachlässigung der Frage der Arterhaltung, Fragen der Methodik der ontogenetischen Forschung, und Aufgaben und Methoden der Evolutionsforschung.

Bei der Einschätzung des Anteils, den Lorenz an der Entwicklung der Ethologie genommen hat und noch nimmt, habe ich als seinen Hauptbeitrag den bezeichnet, daß er uns gezeigt hat, wie man bewährtes “biologisches Denken” folgerichtig auf Verhalten anwenden kann. Daß er dabei an die Arbeit seiner Vorgänger angeknüpft hat, ist nicht mehr verwunderlich, als daß jeder Vater selbst einen Vater hat.

Insbesondere scheint mir das Wesentliche an Lorenz‘ Arbeit zu sein, daß er klar gesehen hat, daß Verhaltensweisen Teile von “Organen”, von Systemen der Arterhaltung sind; daß ihre Verursachung genau so exakt untersucht werden kann wie die gleich welcher anderer Lebensvorgänge, daß ihr arterhaltender Wert ebenso systematisch und exakt aufweisbar ist wie ihre Verursachung, daß Verhaltensontogenie in grundsätzlich gleicher Weise erforscht werden kann wie die Ontogenie der Form und daß die Erforschung der Verhaltensevolution der Untersuchung der Strukturevolution parallel geht. Und obwohl Lorenz ein riesiges Tatsachenmaterial gesammelt hat, ist die Ethologie doch noch mehr durch seine Fragestellung und durch kühne Hypothesen gefördert als durch eigene Nachprüfung dieser Hypothesen. Ohne den Wert solcher Nachprüfung zu unterschätzen — ohne die es natürlich keine Weiterentwicklung gäbe — möchte ich doch behaupten, daß die durch Nachprüfung notwendig gewordenen Modifikationen neben der Leistung des ursprünglichen Ansatzes vergleichsweise unbedeutend sind.

Nebenbei sei auch daran erinnert, daß eine der vielen heilsamen Nachwirkungen der Lorenzschen Arbeit das wachsende Interesse ist, das die Humanpsychologie der Ethologie entgegenbringt ‐ ein erster Ansatz einer Entwicklung, deren Tragweite wir noch kaum übersehen können.

Am Schluß noch eine Bemerkung zur Terminologie. Ich habe hier das Wort “Ethologie” auf einen Riesenkomplex von Wissenschaften angewandt, von denen manche, wie Psychologie und Physiologie, schon längst anerkannte Namen tragen. Das heißt natürlich nicht, daß ich den Namen Ethologie für dieses ganze Gebiet vorschlagen will; das wäre geschichtlich einfach falsch, weil das Wort historisch nur die Arbeit einer kleinen Gruppe von Zoologen kennzeichnet. Der Name ist natürlich gleichgültig; worauf es mir vor allem ankommt, ist darzutun, daß wir das Zusammenwachsen vieler Einzeldisziplinen zu einer vielumfassenden Wissenschaft erleben, für die es nur einen richtigen Namen gibt: “Verhaltensbiologie”. Selbstverständlich ist diese synthetische Entwicklung nicht die Arbeit eines Mannes oder gar die der Ethologen. Sie ist die Folge einer allgemeinen Neigung, Brücken zwischen verwandten Wissenschaften zu schlagen, einer Neigung, die sich in vielen Disziplinen entwickelt hat. Unter den Zoologen ist es Lorenz, der hierzu am meisten beigetragen und zudem manche Nachbardisziplinen stärker beeinflußt hat als irgendein anderer. Ich bin sogar davon überzeugt, daß diese Einwirkungen auf Nachbarwissenschaften noch lange anhalten werden und daß die Verhaltensbiologie erst am Anfang ihrer Ontogenie steht.

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  • , A longitudinal study of adolescents’ judgments of the attractiveness of facial symmetry, averageness and sexual dimorphism, Journal of Evolutionary Psychology, 9, 1, (43), (2011).
  • , Teaching behavioral observation in the 21st century, Zoo Biology, 30, 2, (232-234), (2010).
  • , Review of wallowing in pigs: Description of the behaviour and its motivational basis, Applied Animal Behaviour Science, 132, 1-2, (1), (2011).
  • , Neuronal Nitric Oxide Synthase as a Substrate for the Evolution of Pseudosexual Behaviour in a Parthenogenetic Whiptail Lizard, Journal of Neuroendocrinology, 23, 3, (244-253), (2011).
  • , Contrasting Sociality in Two Widespread, Generalist, Mustelid Genera,MelesandMartes, Mammal Study, 36, 4, (169), (2011).
  • , A Roadmap for Bridging Basic and Applied Research in Forensic Entomology, Annual Review of Entomology, 56, 1, (401), (2011).
  • , Teenage pregnancy and motherhood: How might evolutionary theory inform policy?, Journal of Evolutionary Psychology, 9, 1, (3), (2011).
  • , Genes, hormones, and circuits: An integrative approach to study the evolution of social behavior, Frontiers in Neuroendocrinology, 32, 3, (320), (2011).
  • , Evolutionary Psychology and Feminism, Sex Roles, 64, 9-10, (768), (2011).
  • , Note types and coding in Parid vocalizations: The chick-a-dee call of the boreal chickadee (Poecile hudsonicus), The Journal of the Acoustical Society of America, 129, 5, (3327), (2011).
  • , Hormonal and neurochemical mechanisms of aggression and a new classification of aggressive behavior, Aggression and Violent Behavior, 16, 6, (461), (2011).
  • , Animal Cognition: The next steps, Animal Cognition, 14, 5, (621), (2011).
  • , Evolutionary functions of early social modulation of hypothalamic-pituitary-adrenal axis development in humans, Neuroscience & Biobehavioral Reviews, 35, 7, (1611), (2011).
  • , Evolutionary Theory and the Ultimate–Proximate Distinction in the Human Behavioral Sciences, Perspectives on Psychological Science, 6, 1, (38), (2011).
  • , Using Tinbergen’s Four Questions (Plus One) to Facilitate Evolution Education for Human-Oriented Disciplines, Evolution: Education and Outreach, 4, 1, (107), (2011).
  • , Towards meeting Tinbergen's challenge, Hormones and Behavior, 60, 1, (22), (2011).
  • 2011 IEEE International Conference on Development and Learning (ICDL) Frankfurt am Main, Germany 2011 IEEE International Conference on Development and Learning (ICDL) IEEE , (2011). 978-1-61284-989-8 What are intrinsic motivations? A biological perspective , (2011). 1 8 6037367 , 10.1109/DEVLRN.2011.6037367 http://ieeexplore.ieee.org/lpdocs/epic03/wrapper.htm?arnumber=6037367
  • , Drugs as instruments: A new framework for non-addictive psychoactive drug use, Behavioral and Brain Sciences, 34, 06, (293), (2011).
  • , Oxytocin and social motivation, Developmental Cognitive Neuroscience, 1, 4, (471), (2011).
  • , Logical Justification of the Ethological Study of Play: Ethology Meets Prototype Theory, Primate Research, 27, 2, (127), (2011).
  • , A Population Genetics-Phylogenetics Approach to Inferring Natural Selection in Coding Sequences, PLoS Genetics, 7, 12, (e1002395), (2011).
  • , The Behavioral Ecology of Nutrient Foraging by Plants, Annual Review of Ecology, Evolution, and Systematics, 42, 1, (289), (2011).
  • , Sixteen common misconceptions about the evolution of cooperation in humans, Evolution and Human Behavior, 32, 4, (231), (2011).
  • , Conceptualizing the Autism Spectrum in Terms of Natural Selection and Behavioral Ecology: The Solitary Forager Hypothesis, Evolutionary Psychology, 9, 2, (147470491100900), (2011).
  • , Only full-sibling families evolved eusociality, Nature, 471, 7339, (E4), (2011).
  • , Closing the circle between perceptions and behavior: A cybernetic view of behavior and its consequences for studying motivation and development, Developmental Cognitive Neuroscience, 1, 4, (404), (2011).
  • , Where have all the trees gone? The declining use of phylogenies in animal behaviour journals, Animal Behaviour, 81, 3, (667), (2011).
  • , Reasoning serves argumentation in children, Cognitive Development, 26, 3, (177), (2011).
  • , Associative symmetry in a spatial sample-response paradigm, Behavioural Processes, 86, 3, (305), (2011).
  • , Beyond Generalized Darwinism. II. More Things in Heaven and Earth, Biological Theory, 6, 4, (351), (2011).
  • , Animal models for depression and the mismatch hypothesis of disease, Psychoneuroendocrinology, 36, 3, (330), (2011).
  • , The C. elegans Touch Response Facilitates Escape from Predacious Fungi, Current Biology, 21, 15, (1326), (2011).
  • , The natural order?, Journal of Evolutionary Psychology, 9, 2, (195), (2011).
  • , Developing multiple hypotheses in behavioral ecology, Behavioral Ecology and Sociobiology, 65, 1, (37), (2011).
  • , Existential neuroscience: a proximate explanation of religion as flexible meaning and palliative, Religion, Brain & Behavior, 1, 3, (244), (2011).
  • , Friends with Benefits: The Evolved Psychology of Same- and Opposite-Sex Friendship, Evolutionary Psychology, 9, 4, (147470491100900), (2011).
  • , Sympathy with Adam Smith and reflexions on self, Journal of Economic Behavior & Organization, 77, 1, (4), (2011).
  • , Darwin in Mind: New Opportunities for Evolutionary Psychology, PLoS Biology, 9, 7, (e1001109), (2011).
  • , Book review, Applied Animal Behaviour Science, 131, 3-4, (153), (2011).
  • , Characterization of the Dopamine System in the Brain of the Túngara Frog, Physalaemus pustulosus, Brain, Behavior and Evolution, 10.1159/000321715, 76, 3-4, (211-225), (2010).
  • , Born to Cooperate? Altruism as Exaptation and the Evolution of Human Sociality, Origins of Altruism and Cooperation, 10.1007/978-1-4419-9520-9_4, (41-61), (2011).
  • , Toward a Neuroethology of Schizophrenia: Findings from the Crimean Project, Handbook of Schizophrenia Spectrum Disorders, Volume II, 10.1007/978-94-007-0831-0_6, (121-164), (2011).
  • , The Evolution of Relevance, Cognitive Science, 34, 4, (583-601), (2010).
  • , The relationship between leadership and gregariousness in mixed‐species bird flocks, Journal of Avian Biology, 41, 1, (99-103), (2010).
  • , Adaptation and the Evolution of Physiological Characters, Comprehensive Physiology, (3-16), (2011).
  • , Comparative Psychology, The Corsini Encyclopedia of Psychology, (1-3), (2010).
  • , Development and Evolution Revisited, Handbook of Developmental Science, Behavior, and Genetics, (61-109), (2010).
  • , Action in Cognitive Ethology, A Companion to the Philosophy of Action, (393-400), (2010).
  • , Evolutionary psychology and the origins of language, Journal of Evolutionary Psychology, 8, 4, (289), (2010).
  • , ‘Birdwatching and baby-watching’: Niko and Elisabeth Tinbergen’s ethological approach to autism, History of Psychiatry, 21, 2, (176), (2010).
  • , Why Are There Social Gradients in Preventative Health Behavior? A Perspective from Behavioral Ecology, PLoS ONE, 5, 10, (e13371), (2010).
  • , The Evolutionary Origins of Human Generosity, International Sociology, 25, 3, (443), (2010).
  • , Heart Rate during Conflicts Predicts Post-Conflict Stress-Related Behavior in Greylag Geese, PLoS ONE, 5, 12, (e15751), (2010).
  • , The Four Whys of Age‐Linked Regression Periods in Infancy, Nurturing Children and Families, (107-119), (2010).
  • , Nick Thompson on Design, Integrative Psychological and Behavioral Science, 44, 4, (351), (2010).
  • , A review and tests of hypotheses about causes of the KwaZulu-Natal sardine run, African Journal of Marine Science, 32, 2, (449), (2010).
  • , Genetic Contributions to Antisocial Personality and Behavior: A Meta-Analytic Review From an Evolutionary Perspective, The Journal of Social Psychology, 150, 2, (160), (2010).
  • , The evolutionary turn in psychiatry: a historical overview, History of Psychiatry, 21, 2, (131), (2010).
  • , Encyclopedia of Animal Behavior, (428), (2010).
  • , The Informal Corporate Identity Communication Process, Corporate Reputation Review, 13, 3, (157), (2010).
  • , The proximate mechanisms and ultimate functions of smiles, Behavioral and Brain Sciences, 33, 06, (454), (2010).
  • , The Nature of Neuroethology, Brain, Behavior and Evolution, 76, 3-4, (163), (2010).
  • , Further Thoughts on the Evolution of Pride’s Two Facets: A Response to Clark, Emotion Review, 2, 4, (399), (2010).
  • , Integration of Proximate and Ultimate Causes, Encyclopedia of Animal Behavior, 10.1016/B978-0-08-045337-8.00207-2, (180-185), (2010).
  • , Absolute pitch in boreal chickadees and humans: Exceptions that test a phylogenetic rule, Learning and Motivation, 41, 3, (156), (2010).
  • , Book review, Physiology & Behavior, 101, 3, (401), (2010).
  • , Developmental stress and female mate choice behaviour in the zebra finch, Animal Behaviour, 79, 6, (1381), (2010).
  • , Size-related habitat shifts facilitated by positive preference induction in a marine kelp crab, Behavioral Ecology, 21, 2, (329), (2010).
  • , Book Review: Climbing Mount Impossible: An Important Book Written Too EarlyA review of BrüneMartin, Textbook of Evolutionary Psychiatry: The Origins of Psychopathology. Oxford University Press: New York, 2008, 385 pp., US$67.95, ISBN 978-0-19-920768-8 (paperback)., Evolutionary Psychology, 8, 2, (147470491000800), (2010).
  • , Maladaptive behavior reinforces a recruitment bottleneck in newly settled fishes, Oecologia, 164, 1, (99), (2010).
  • , Current Opinion in Neurobiology - Sensory systems, Current Opinion in Neurobiology, 20, 3, (271), (2010).
  • , Renovating the Pyramid of Needs, Perspectives on Psychological Science, 5, 3, (292), (2010).
  • , Encyclopedia of Animal Behavior, (340), (2010).
  • , Encyclopedia of Animal Behavior, (335), (2010).
  • , A Naturalist’s View of Pride, Emotion Review, 2, 2, (163), (2010).
  • , Evolutionary explanations in medicine: How do they differ and how to benefit from them, Medical Hypotheses, 74, 4, (746), (2010).
  • , Tonic Communication in the Antipredator Behavior of Ground Squirrels, , 10.1016/S0065-3454(10)41004-9, (119-149), (2010).
  • , ROBERT C. RICHARDSON * Evolutionary Psychology as Maladapted Psychology, The British Journal for the Philosophy of Science, 61, 4, (883), (2010).
  • , Levels of Explanation in Behavioral Ecology, California Archaeology, 2, 1, (77), (2010).
  • , Encyclopedia of Animal Behavior, (649), (2010).
  • , Microevolution and Macroevolution in Behavior, Encyclopedia of Animal Behavior, 10.1016/B978-0-08-045337-8.00114-5, (449-454), (2010).
  • , Toward an Integrative Approach of Cognitive Neuroscientific and Evolutionary Psychological Studies of Art, Evolutionary Psychology, 8, 4, (147470491000800), (2010).
  • , Putting evolution back into the social sciences, Journal of Evolutionary Psychology, 8, 2, (127), (2010).
  • , Encyclopedia of Animal Behavior, (557), (2010).
  • , Darwin's Rehabilitation of Teleology Versus Williams' Replacement of Teleology by Natural Selection, Biological Theory, 5, 4, (357), (2010).
  • , Evolving the Capacity to Understand Actions, Intentions, and Goals, Annual Review of Psychology, 61, 1, (303), (2010).
  • , Individual performance in complex social systems: the greylag goose example, Animal Behaviour: Evolution and Mechanisms, 10.1007/978-3-642-02624-9_5, (121-148), (2010).
  • , The lost dream of ecological determinism: Time to say goodbye? … Or a White Queen's proposal?, "Evolutionary Anthropology: Issues, News, and Reviews", 18, 5, (166-174), (2009).
  • , Integration of proximate and evolutionary explanation of reproductive strategy: The case of callitrichid primates and implications for human biology, American Journal of Human Biology, 21, 6, (731-738), (2009).
  • , The ecology and evolutionary endocrinology of reproduction in the human female, American Journal of Physical Anthropology, 140, S49, (95-136), (2009).
  • , The study of behaviour in personality psychology: Meaning, importance and measurement, European Journal of Personality, 23, 5, (437-453), (2009).
  • , Nutrition, ecology and nutritional ecology: toward an integrated framework, Functional Ecology, 23, 1, (4-16), (2009).
  • , Ethics and Animal Consciousness: How Rubber the Ethical Ruler?, Journal of Social Issues, 65, 3, (499-521), (2009).
  • , Evolution in anthropology: a comment on Nettle's ‘Beyond nature versus culture’, Journal of the Royal Anthropological Institute, 15, 2, (244-246), (2009).
  • , The evolution of comparative cognition: Is the snark still a boojum?, Behavioural Processes, 80, 3, (210), (2009).
  • , Depression in adults: Some basic facts, Psychoanalytic Psychotherapy, 23, 3, (225), (2009).
  • , Evolutionary approach to communication between humans and dogs, Veterinary Research Communications, 33, S1, (53), (2009).
  • , The socio-relational framework of expressive behaviors as an integrative psychological paradigm, Behavioral and Brain Sciences, 32, 05, (408), (2009).
  • , The Vernacular Concept of Innateness, Mind & Language, 24, 5, (605-630), (2009).
  • , Four noble questions, Ethology, 115, 8, (798-799), (2009).
  • , Experimental Semiotics: A New Approach for Studying Communication as a Form of Joint Action, Topics in Cognitive Science, 1, 2, (393-410), (2009).
  • , The roots of human altruism, British Journal of Psychology, 100, 3, (455-471), (2011).
  • , In Defense of the Abstracted Animat, Adaptive Behavior, 17, 4, (303), (2009).
  • , , Lingua, 119, 7, (1106), (2009).
  • , Psychology and the grand unified theory, Journal of Evolutionary Psychology, 7, 4, (359), (2009).
  • , In What Sense Does ‘Nothing Make Sense Except in the Light of Evolution’?, Acta Biotheoretica, 57, 1-2, (11), (2009).
  • , A Melding of the Minds: When Primatology Meets Personality and Social Psychology, Personality and Social Psychology Review, 13, 2, (129), (2009).
  • , A designer's view: The perspective of form and function, Journal of Biological Education, 44, 1, (12), (2009).
  • , Artificial Immune Systems, (22)
  • , Marital status, gender and cardiovascular mortality: Behavioural, psychological distress and metabolic explanations, Social Science & Medicine, 69, 2, (223), (2009).
  • , Focusing the metaphor: plant root foraging behaviour, Trends in Ecology & Evolution, 24, 8, (419), (2009).
  • , Plant intelligence, Plant Signaling & Behavior, 4, 5, (394), (2009).
  • , Dyadic relationships and operational performance of male and female owners and their male dogs, Behavioural Processes, 81, 3, (383), (2009).
  • , The synthetic approach to the study of spatial memory: Have we properly addressed Tinbergen's “four questions”?, Behavioural Processes, 80, 3, (278), (2009).
  • , Current status and future directions of research on facial attractiveness, Anthropological Review, 72, 1, (2009).
  • , The nature of human aggression, International Journal of Law and Psychiatry, 32, 4, (202), (2009).
  • , Niko Tinbergen and the red patch on the herring gull's beak, Animal Behaviour, 77, 4, (785), (2009).
  • , Ethological approaches to psychiatric disorders: focus on depression and schizophrenia, Australian and New Zealand Journal of Psychiatry, 43, 11, (1007), (2009).
  • , Love thy neighbour? Social nesting pattern, host mass and nest size affect ectoparasite intensity in Darwin’s tree finches, Behavioral Ecology and Sociobiology, 63, 5, (731), (2009).
  • , Chapter 5 Behavior of Fishes in the Sexual/Unisexual Mating System of the Amazon Molly (Poecilia formosa), , 10.1016/S0065-3454(09)39005-1, (153-183), (2009).
  • , Dead or Alive? Knowledge about a Sibling's Death Varies by Genetic Relatedness in a Modern Society, Evolutionary Psychology, 7, 1, (147470490900700), (2009).
  • , Sind psychische Störungen etwas spezifisch Menschliches?, Der Nervenarzt, 80, 3, (252), (2009).
  • , Refining the sexual selection explanation within an ethological framework, Behavioral and Brain Sciences, 32, 3-4, (292), (2009).
  • , Comparative Psychology as an Effective Supplement to Undergraduate Core Psychology Courses, Teaching of Psychology, 36, 3, (200), (2009).
  • , Sequential and simultaneous choices: Testing the diet selection and sequential choice models, Behavioural Processes, 80, 3, (218), (2009).
  • , Animal behaviour, animal welfare and the scientific study of affect, Applied Animal Behaviour Science, 118, 3-4, (108), (2009).
  • , Life history as an integrative theoretical framework advancing the understanding of the attachment system, Behavioral and Brain Sciences, 32, 01, (34), (2009).
  • , What can function tell us about mechanism?, Trends in Ecology & Evolution, 24, 7, (357), (2009).
  • , Sexual Coercion in Intimate Relationships: A Comparative Analysis of the Effects of Women’s Infidelity and Men’s Dominance and Control, Archives of Sexual Behavior, 38, 2, (226), (2009).
  • , The evolution of combinatorial phonology, Journal of Phonetics, 37, 2, (125), (2009).
  • , Chapter 3 The Dog as a Model for Understanding Human Social Behavior, , 10.1016/S0065-3454(09)39003-8, (71-116), (2009).
  • , Dossier Évolution et créationnisme. Dépasser les oppositions et aborder la téléologie en questionnant la notion de vie, Natures Sciences Sociétés, 17, 1, (50), (2009).
  • , Behavioural biologists do not agree on what constitutes behaviour, Animal Behaviour, 78, 1, (103), (2009).
  • , On the implications of the classical ergodic theorems: Analysis of developmental processes has to focus on intra‐individual variation, Developmental Psychobiology, 50, 1, (60-69), (2007).
  • , Three methodological core issues of comparative personality research, European Journal of Personality, 22, 5, (475-496), (2008).
  • , The Science of Animal Suffering, Ethology, 114, 10, (937-945), (2008).
  • , Distinguishing four fundamental approaches to the evolution of helping, Journal of Evolutionary Biology, 21, 2, (405-420), (2008).
  • , The great opportunity: Evolutionary applications to medicine and public health, Evolutionary Applications, 1, 1, (28-48), (2008).
  • , The Use of Theory in Behavioural Research, Ethology, 114, 1, (1-6), (2008).
  • , Comparative Psychology, Handbook of Research Methods in Experimental Psychology, (209-240), (2008).
  • , Cognitive Models of Human–Information Interaction, Handbook of Applied Cognition, (443-470), (2008).
  • , A Bayesian Approach to Optimal Foraging in Parasitoids, Behavioral Ecology of Insect Parasitoids, (357-383), (2008).
  • , The Question of Animal Emotions: An Ethological Perspective, Mental Health and Well‐Being in Animals, (15-27), (2008).
  • , A Neuroethological Perspective on the Perception of Vocal Communication Signals, The Handbook of Speech Perception, (653-675), (2008).
  • , Cognitive Ethology for humans: Inconvenient truth or attentional deficit?, British Journal of Psychology, 99, 3, (347-350), (2010).
  • , The Role of Development on Evolutionary Psychology: Tinbergen Revisited, Psychological Inquiry, 19, 1, (38), (2008).
  • , Evidence for conditional sex differences in emotional but not in sexual jealousy at the automatic level of cognitive processing, European Journal of Personality, 22, 1, (3-30), (2007).
  • , Infant mortality following male takeovers in wild geladas, American Journal of Primatology, 70, 12, (1152-1159), (2008).
  • , Function and Teleology, A Companion to the Philosophy of Biology, (525-549), (2008).
  • , Ethology, Sociobiology, and Evolutionary Psychology, A Companion to the Philosophy of Biology, (393-414), (2008).
  • , How Producer Biases Can Favor the Evolution of Communication: An Analysis of Evolutionary Dynamics, Adaptive Behavior, 16, 1, (27), (2008).
  • , Integrating perception and action through cognitive neuropsychology (broadly conceived), Cognitive Neuropsychology, 25, 7-8, (879), (2008).
  • , Epigenetics and its implications for behavioral neuroendocrinology, Frontiers in Neuroendocrinology, 29, 3, (344), (2008).
  • , Complex display behaviour ofEvarcha culicivora, an East African mosquito‐eating jumping spider, New Zealand Journal of Zoology, 35, 2, (151), (2008).
  • , Species-Specific Diversity of a Fixed Motor Pattern: The Electric Organ Discharge of Gymnotus, PLoS ONE, 3, 5, (e2038), (2008).
  • , Adaptive learning by a target‐tracking system, International Journal of Intelligent Computing and Cybernetics, 1, 1, (46), (2008).
  • , Chapter 1 Using Robots to Understand Animal Behavior, , 10.1016/S0065-3454(08)00001-6, (1-58), (2008).
  • , Beyond persons: extending the personal/subpersonal distinction to non-rational animals and artificial agents, Biology & Philosophy, 23, 1, (87), (2008).
  • , New Zealand trends in animal behaviour research, New Zealand Journal of Zoology, 35, 4, (305), (2008).
  • , Ethics, evolution and culture, Theory in Biosciences, 127, 3, (229), (2008).
  • , Proximate and ultimate explanations are required for a comprehensive understanding of partner rape, Aggression and Violent Behavior, 13, 2, (119), (2008).
  • , The other cooperation problem: generating benefit, Biology & Philosophy, 23, 2, (179), (2008).
  • , Attachment—Past and Present. But What About the Future?, Integrative Psychological and Behavioral Science, 42, 4, (406), (2008).
  • , Short-distance navigation in cephalopods: a review and synthesis, Cognitive Processing, 9, 4, (239), (2008).
  • , Putting the Altruism Back into Altruism: The Evolution of Empathy, Annual Review of Psychology, 59, 1, (279), (2008).
  • , Book Review, Animal Behaviour, 75, 2, (739), (2008).
  • , Comprendre la dépression : apport de la psychologie évolutionniste, PSN, 6, 4, (188), (2008).
  • , Innateness, Canalization, and ‘Biologicizing the Mind’, Philosophical Psychology, 21, 3, (397), (2008).
  • , Learning and Memory in Communication and Navigation in Insects, Learning and Memory: A Comprehensive Reference, 10.1016/B978-012370509-9.00085-1, (477-498), (2008).
  • , A toolkit for stimulating productive thinking, Journal of Biological Education, 43, 1, (21), (2008).
  • , The Debate on Behavior in Conservation: New Zealand Integrates Theory with Practice, BioScience, 58, 5, (454), (2008).
  • , INFLUENCE OF TEMPERATURE ON SWARMBOTS THAT LEARN, Cybernetics and Systems, 39, 5, (502), (2008).
  • , Cooperative Behaviour in Fishes, Fish Behaviour, 10.1201/b10757-16, (437-484), (2014).
  • , Behavioral genetics and development: Historical and conceptual causes of controversy, New Ideas in Psychology, 26, 3, (332), (2008).
  • , Lateralización cerebral en chimpancés: una aproximación filogenética al estudio del cerebro humano, Estudios de Psicología, 10.1174/021093908784485084, 29, 2, (147-161), (2014).
  • , Developing human nature: “Development to” versus “Development from?”, Developmental Psychobiology, 49, 8, (788-799), (2007).
  • , Developing a framework for development: A discussion, Developmental Psychobiology, 49, 1, (77-86), (2006).
  • , The adaptive nature of non‐food collection for the Florida harvester ant, Pogonomyrmex badius, Ecological Entomology, 32, 1, (105-112), (2007).
  • , Social semantics: altruism, cooperation, mutualism, strong reciprocity and group selection, Journal of Evolutionary Biology, 20, 2, (415-432), (2006).
  • , Evolutionary Adaptationism: Another Biological Approach to Criminal and Antisocial Behaviour, Ciba Foundation Symposium 194 ‐ Genetics of Criminal and Antisocial Behaviour, (183-201), (2007).
  • 2007 Frontiers in the Convergence of Bioscience and Information Technologies, (2007).Biologically-Inspired Adaptive Learning: A Near Set Approach40310.1109/FBIT.2007.39
  • , Coexistence in Female‐Bonded Primate Groups, , 10.1016/S0065-3454(07)37002-2, (43-81), (2007).
  • , How Elephants are Opening Doors: Developmental Neuroethology, Attachment and Social Context, Ethology, 113, 5, (426-436), (2007).
  • , Evolutionary explanations, Philosophy of Anthropology and Sociology, 10.1016/B978-044451542-1/50008-8, (249-310), (2007).
  • , The Transition to Experiencing: II. The Evolution of Associative Learning Based on Feelings, Biological Theory, 2, 3, (231), (2007).
  • , Modeling birdsong learning with a chaotic Elman network, Artificial Life and Robotics, 11, 2, (162), (2007).
  • , A family reunion among estranged cousins Review of Primate Psychology edited by Dario Maestripieri. Cambridge, Massachusetts, Harvard University Press, 2003, ix+619 pp, 43 figures, 12 tables., American Journal of Primatology, 69, 3, (353-357), (2007).
  • , The evolution of IT innovations in Swedish organizations: a Darwinian critique of ‘Lamarckian’ institutional economics, Journal of Evolutionary Economics, 17, 1, (1), (2007).
  • , Hormones and the development of sex differences in behavior, Journal of Ornithology, 10.1007/s10336-007-0188-3, 148, S1, (17-26), (2007).
  • , Engineering Human Cooperation, Human Nature, 18, 2, (88), (2007).
  • , Sex and species differences in spatial memory in food-storing kangaroo rats, Animal Behaviour, 73, 2, (321), (2007).
  • , Evolutionary Explanations for Cooperation, Current Biology, 17, 16, (R661), (2007).
  • , The Evolution, Function, and Meaning of Marmot Alarm Communication, , 10.1016/S0065-3454(07)37008-3, (371-401), (2007).
  • , Translocation reverses birth sex ratio bias depending on its timing during gestation: evidence for the action of two sex-allocation mechanisms, Reproduction, Fertility and Development, 19, 7, (831), (2007).
  • , Equine learning in a wider context—Opportunities for integrative pluralism, Behavioural Processes, 76, 1, (53), (2007).
  • , The Social Evolution of Language, and the Language of Social Evolution, Evolutionary Psychology, 5, 4, (147470490700500), (2007).
  • , Developmental environment, cultural transmission, and mate choice copying, Naturwissenschaften, 94, 8, (651), (2007).
  • , Evaluating methods to quantify anthropogenic stressors on wild animals, Applied Animal Behaviour Science, 102, 3-4, (429), (2007).
  • , Where Evolutionary Psychology Meets Cognitive Neuroscience: A Précis to Evolutionary Cognitive Neuroscience, Evolutionary Psychology, 5, 1, (147470490700500), (2007).
  • , Neo‐functional Analysis: Phylogenetical Restrictions on Causal Role Functions, Philosophy of Science, 74, 5, (601), (2007).
  • , History of ethology comes of age, Biology & Philosophy, 23, 1, (129), (2007).
  • 2007 IEEE Symposium on Foundations of Computational Intelligence, (2007).Cassifctinof Objects by-Means of Features29610.1109/FOCI.2007.372183
  • , Approximation spaces in off-policy Monte Carlo learning, Engineering Applications of Artificial Intelligence, 20, 5, (667), (2007).
  • 2007 IEEE Symposium on Foundations of Computational Intelligence, (2007).Granular Computing in Actor-Critic Learning5910.1109/FOCI.2007.372148
  • , Campaign Effects and Second-Order Cycles, European Union Politics, 8, 4, (509), (2007).
  • , Niko Tinbergen: The Ethologist as Field Naturalist, Biological Theory, 2, 1, (87), (2007).
  • , A gene mutation which changed animal behaviour: Margaret Bastock and the yellow fly, Animal Behaviour, 74, 2, (163), (2007).
  • , Behavioural biology: an effective and relevant conservation tool, Trends in Ecology & Evolution, 22, 8, (401), (2007).
  • , Phylogenetic analysis and alignment of behavioral sequences by direct optimization, Cladistics, 22, 6, (602-633), (2006).
  • , A bias for social information in human cultural transmission, British Journal of Psychology, 97, 3, (405-423), (2010).
  • , The Multipredator Hypothesis and the Evolutionary Persistence of Antipredator Behavior, Ethology, 112, 3, (209-217), (2006).
  • , The Rise and Fall of Ethology, Ethology, 112, 7, (726-728), (2006).
  • , An accessible introduction to animal behaviour, Global Ecology and Biogeography, 15, 1, (109-110), (2006).
  • , Sociobiology, Encyclopedia of Cognitive Science, (2006).
  • , A comparative analysis of animals' understanding of the human pointing gesture, Animal Cognition, 9, 2, (81), (2006).
  • , Ethology into a new era, Ethology, 112, 1, (1-6), (2006).
  • , Introduction, Ornithological Science, 5, 1, (1), (2006).
  • , The Zebrafish as a Model for Behavioral Studies, Zebrafish, 3, 2, (227), (2006).
  • , Sexual Reward via Vulvar, Perineal, and Anal Stimulation: A Proximate Mechanism for Female Homosexual Mounting in Japanese Macaques, Archives of Sexual Behavior, 35, 5, (523), (2006).
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  • , Primate group size, brains and communication: A New World perspective, Behavioral and Brain Sciences, 16, 04, (711), (1993).
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  • , Vocal grooming: Man the schmoozer, Behavioral and Brain Sciences, 16, 04, (699), (1993).
  • , Size of human groups during the Paleolithic and the evolutionary significance of increased group size, Behavioral and Brain Sciences, 16, 04, (709), (1993).
  • , Grooming is not the only regulator of primate social interactions, Behavioral and Brain Sciences, 16, 04, (717), (1993).
  • , Brains, grouping and language, Behavioral and Brain Sciences, 16, 04, (706), (1993).
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  • , Sizing up social groups, Behavioral and Brain Sciences, 16, 04, (710), (1993).
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  • , Anthropological criticisms of Dunbar's theory of the origin of language, Behavioral and Brain Sciences, 16, 04, (705), (1993).
  • , The rest of the story: Grooming, group size and vocal exchanges in neotropical primates, Behavioral and Brain Sciences, 16, 04, (718), (1993).
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  • , Do larger brains mean greater intelligence?, Behavioral and Brain Sciences, 16, 04, (696), (1993).
  • , Comparative studies, phylogenies and predictions of coevolutionary relationships, Behavioral and Brain Sciences, 16, 04, (714), (1993).
  • , Another primate brain fiction: Brain (cortex) weight and homogeneity, Behavioral and Brain Sciences, 16, 04, (707), (1993).
  • , Group structure and group size among humans and other primates, Behavioral and Brain Sciences, 16, 04, (703), (1993).
  • , Language and levels of selection, Behavioral and Brain Sciences, 16, 04, (701), (1993).
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  • , Number our days: Quantifying social evolution, Behavioral and Brain Sciences, 16, 04, (712), (1993).
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  • , A developmental look at grooming, grunting and group cohesion, Behavioral and Brain Sciences, 16, 04, (716), (1993).
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  • , Essay on Contemporary Issues in Ethology: On the Problems Studied in Ethology, Comparative Psychology, and Animal Behavior, Ethology, 92, 2, (89-107), (2010).
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  • , ON IMMEDIATE FUNCTION, Journal of the Experimental Analysis of Behavior, 57, 3, (417-427), (2013).
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  • , Calls as labels: An intriguing theme, but one with limitations, Behavioral and Brain Sciences, 15, 01, (162), (1992).
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  • , What are mental states?, Behavioral and Brain Sciences, 15, 01, (162), (1992).
  • , Characterizing the mind of another species, Behavioral and Brain Sciences, 15, 01, (172), (1992).
  • , Is lack of understanding of cause-effect relationships a suitable basis for interpreting monkeys' failures in attribution?, Behavioral and Brain Sciences, 15, 01, (169), (1992).
  • , Looking inside monkey minds: Milestone or millstone, Behavioral and Brain Sciences, 15, 01, (150), (1992).
  • , Behavioral Responses of Insects to Plant Secondary Compounds, Herbivores: Their Interactions with Secondary Plant Metabolites, 10.1016/B978-0-08-092545-5.50007-9, (45-88), (1992).
  • , Animal mentality: Canons to the right of them, canons to the left of them …, Behavioral and Brain Sciences, 15, 01, (154), (1992).
  • , Monkeys mind, Behavioral and Brain Sciences, 15, 01, (147), (1992).
  • , Knowing thyself, knowing the other: They're not the same, Behavioral and Brain Sciences, 15, 01, (166), (1992).
  • , Précis of How monkeys see the world, Behavioral and Brain Sciences, 15, 01, (135), (1992).
  • , Mind reading, pretence and imitation in monkeys and apes, Behavioral and Brain Sciences, 15, 01, (170), (1992).
  • , Theory of society, yes, theory of mind, no, Behavioral and Brain Sciences, 15, 01, (155), (1992).
  • , Are monkeys nomothetic or idiographic?, Behavioral and Brain Sciences, 15, 01, (161), (1992).
  • , Is the monkeys' world scientifically impenetrable?, Behavioral and Brain Sciences, 15, 01, (152), (1992).
  • , Exploring the “boundary” between the minds of monkeys and humans, Behavioral and Brain Sciences, 15, 01, (163), (1992).
  • , Monkeys and consciousness, Behavioral and Brain Sciences, 15, 01, (147), (1992).
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  • , HOW DID MORALITY EVOLVE?, Zygon®, 26, 1, (49-89), (2005).
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  • , Too many errors, Behavioral and Brain Sciences, 14, 02, (306), (1991).
  • , When the mind goes awry: Schizophrenia and the emergence of culture, Behavioral and Brain Sciences, 14, 02, (307), (1991).
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  • , Organizational levels analysis: A key to understanding processes in natural systems, Journal of Theoretical Biology, 149, 2, (203), (1991).
  • , Hostile aggression as social skills deficit or evolutionary strategy?, Behavioral and Brain Sciences, 14, 02, (315), (1991).
  • , Freud and sociobiology, Behavioral and Brain Sciences, 14, 02, (319), (1991).
  • , Multiple causes of human behavior, Behavioral and Brain Sciences, 14, 02, (313), (1991).
  • , Person schemas: Evolutionary, individual developmental and social sources, Behavioral and Brain Sciences, 14, 02, (309), (1991).
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  • , Précis of Darwin, sex and status: Biological approaches to mind and culture, Behavioral and Brain Sciences, 14, 02, (295), (1991).
  • , Being aware of consciousness and cultures, Behavioral and Brain Sciences, 14, 02, (316), (1991).
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  • , Once more with feeling: Genes, mind and culture, Behavioral and Brain Sciences, 14, 02, (317), (1991).
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  • , The awakened brain: From Wright's psychozoology to Barkow's selfless persons, Behavioral and Brain Sciences, 14, 02, (311), (1991).
  • , Human nature and the Holy Grail, Behavioral and Brain Sciences, 14, 02, (312), (1991).
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  • , Causes, proximate and ultimate, Biology & Philosophy, 5, 4, (401), (1990).
  • , The Evolutionary Functions of Repression and the Ego Defenses, Journal of the American Academy of Psychoanalysis, 18, 2, (260), (1990).
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  • , Rethinking human ethology, Ethology and Sociobiology, 10, 5, (315), (1989).
  • , Similarity and ethnicity mediate human relationships, but why?, Behavioral and Brain Sciences, 12, 03, (548), (1989).
  • , Ontogenèse et phylogenèse en éthologie, Geobios, 22, (99), (1989).
  • , Evolutionary patterns of host utilization by ichneumonoid parasitoids (Hymenoptera: Ichneumonidae and Braconidae)*, Biological Journal of the Linnean Society, 35, 4, (351-377), (2008).
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  • , The ethologic model of phonetic development: I. Phonetic contrast estimators, Clinical Linguistics & Phonetics, 2, 4, (347), (1988).
  • , Nursing Research and the Discipline of Ethological Science, Western Journal of Nursing Research, 10, 6, (743), (1988).
  • , The levels of analysis, Animal Behaviour, 36, 2, (616), (1988).
  • , An ethologist's approach to behavioural toxicology, Neurotoxicology and Teratology, 10, 2, (85), (1988).
  • , Human cyclic motility: Fetal–Newborn continuities and newborn state differences, Developmental Psychobiology, 20, 4, (425-442), (2004).
  • , “Scotch'd the snake, not killed it”, Behavioral and Brain Sciences, 10, 01, (83), (1987).
  • , Leapfrog over the brain, Behavioral and Brain Sciences, 10, 01, (73), (1987).
  • , The hypothalamus and the impartial perspective, Behavioral and Brain Sciences, 10, 01, (84), (1987).
  • , Testing sociobiological hypotheses ethnographically, Behavioral and Brain Sciences, 10, 01, (74), (1987).
  • , Deception and Evolutionary Biology, Cultural Anthropology, 2, 1, (50-64), (2009).
  • , The development of a developmentalist: Daniel S. Lehrman, Developmental Psychobiology, 20, 6, (563-570), (2004).
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  • , The problem of adaptation in the study of human behavior, Ethology and Sociobiology, 8, 1, (61), (1987).
  • , Confessions of a curmudgeon, Behavioral and Brain Sciences, 10, 01, (89), (1987).
  • , Criticism and realism, Behavioral and Brain Sciences, 10, 01, (72), (1987).
  • , An Ethological Perspective on Homosexuality during Adolescence, Journal of Adolescent Research, 2, 3, (283), (1987).
  • , Saving sociobiology: The use and abuse of logic, Behavioral and Brain Sciences, 10, 01, (73), (1987).
  • , Pop sociobiology and meta-ethics, Behavioral and Brain Sciences, 10, 01, (83), (1987).
  • , Rising out of the ashes, Behavioral and Brain Sciences, 10, 01, (79), (1987).
  • , Familiarity out-breeds, Behavioral and Brain Sciences, 10, 01, (71), (1987).
  • , Useful distinctions in human sociobiology, Behavioral and Brain Sciences, 10, 01, (79), (1987).
  • , Ontogenetic Development of Behavior: The Cricket Visual World, , 10.1016/S0065-3454(08)60179-5, (165-212), (1987).
  • , Darwin and human nature, Behavioral and Brain Sciences, 10, 01, (89), (1987).
  • , Enough of polemics – let's look at data!, Behavioral and Brain Sciences, 10, 01, (79), (1987).
  • , Précis of Vaulting Ambition: Sociobiology and the Quest for Human Nature, Behavioral and Brain Sciences, 10, 01, (61), (1987).
  • , Faulting ambition: A double standard?, Behavioral and Brain Sciences, 10, 01, (78), (1987).
  • , Optimist/pessimist, Behavioral and Brain Sciences, 10, 01, (87), (1987).
  • , Species are individuals: Therefore human nature is a metaphysical delusion, Behavioral and Brain Sciences, 10, 01, (77), (1987).
  • , Putting sociobiology in its place, Behavioral and Brain Sciences, 10, 01, (76), (1987).
  • , Folk psychology versus pop sociobiology, Behavioral and Brain Sciences, 10, 01, (85), (1987).
  • , Bridging the sociobiological gap, Behavioral and Brain Sciences, 10, 01, (88), (1987).
  • , Amplifying sociobiology's hollow ring, Behavioral and Brain Sciences, 10, 01, (78), (1987).
  • , Is human sociobiology a progressive or a degenerating research programme?, Behavioral and Brain Sciences, 10, 01, (86), (1987).
  • , Human communication and contemporary evolutionary theory, Research on Language & Social Interaction, 20, 1-4, (79), (1987).
  • , Sociobiology and the problem of culture, Behavioral and Brain Sciences, 10, 01, (75), (1987).
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  • , Breeding experience and parental behaviour in convict cichlids (), Behavioural Processes, 13, 1-2, (101), (1986).
  • , Chemical communication in hermaphroditic digenetic trematodes, Journal of Chemical Ecology, 12, 8, (1659), (1986).
  • , Do honeybees get tired? The effect of load weight on patch departure, Animal Behaviour, 34, 4, (1243), (1986).
  • , When to feed: decision making in sticklebacks, Gasterosteus aculeatus, Environmental Biology of Fishes, 16, 1-3, (95), (1986).
  • , The making of ethology: The association for the Study of Animal Behaviour, 1936–1986, Animal Behaviour, 34, 6, (1601), (1986).
  • , Motor Patterns in Development, Developmental Psychobiology and Developmental Neurobiology, 10.1007/978-1-4613-2113-2_2, (35-97), (1986).
  • , Functional hypotheses and their impact on behavioral description, Behavioral and Brain Sciences, 8, 01, (187), (1985).
  • , Vocalizations of one-year-olds, Journal of Child Language, 12, 03, (1985).
  • , Directive use in a migrant agricultural community: A test of Ervin-Tripp's hypotheses, Language in Society, 14, 01, (63), (1985).
  • , Behavioral description and its impact on functional inference, Behavioral and Brain Sciences, 8, 01, (186), (1985).
  • , La influencia del entorno en la educación: la aportación de los modelos ecológicos, Infancia y Aprendizaje, 8, 29, (3), (1985).
  • , Benzodiazepines and their antagonists: A pharmacoethological analysis with particular reference to effects on “aggression”, Neuroscience & Biobehavioral Reviews, 9, 1, (21), (1985).
  • , Ethological and sociobiological approaches to the study of the adaptation: Implications for the study of agonistic behavior in primates, Ethology and Sociobiology, 5, 4, (221), (1984).
  • , The biological function of Pavlovian conditioning: Learned inhibition of aggressive behavior in territorial fish, Learning and Motivation, 15, 4, (459), (1984).
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  • , The Case for Praxics, The Behavior Analyst, 7, 2, (101), (1984).
  • , Biological constraints on instrumental and classical conditioning: Retrospect and prospect, Animal Learning & Behavior, 11, 2, (151), (1983).
  • , Awareness and Griffin's circular reasoning, Animal Behaviour, 31, 2, (613), (1983).
  • , Predatory Strategies and Behavioral Diversity, The American Biology Teacher, 45, 6, (334), (1983).
  • , Development of Hand-Use Preference during Infancy, Manual Specialization and the Developing Brain, 10.1016/B978-0-12-773140-7.50007-8, (33-70), (1983).
  • , Biological Constraints and the Pursuit of General Theories of Learning, Animal Cognition and Behavior, 10.1016/S0166-4115(08)61801-4, (319-344), (1983).
  • , Functional aspects of play as revealed by structural components and social interaction patterns, Behavioral and Brain Sciences, 5, 01, (156), (1982).
  • , Play as whimsy, Behavioral and Brain Sciences, 5, 01, (166), (1982).
  • , Play: Structure and function, Behavioral and Brain Sciences, 5, 01, (162), (1982).
  • , The nature-nurture error again, Behavioral and Brain Sciences, 5, 01, (155), (1982).
  • , The essentials of play?, Behavioral and Brain Sciences, 5, 01, (171), (1982).
  • , Play as a mode, Behavioral and Brain Sciences, 5, 01, (168), (1982).
  • , Some missed opportunities in theories of play, Behavioral and Brain Sciences, 5, 01, (165), (1982).
  • , Skill and flexibility in animal play behavior, Behavioral and Brain Sciences, 5, 01, (162), (1982).
  • , Pavlovian Conditioning of Signal-Centered Action Patterns and Autonomic Behavior: A Biological Analysis of Function, , 10.1016/S0065-3454(08)60045-5, (1-64), (1982).
  • , The epistemology of the play theorist, Behavioral and Brain Sciences, 5, 01, (170), (1982).
  • , Does being human matter? On some interpretive problems of comparative ludology, Behavioral and Brain Sciences, 5, 01, (160), (1982).
  • , On the evolution of play by means of artificial selection, Behavioral and Brain Sciences, 5, 01, (165), (1982).
  • , Play—immediate or long-term adaptiveness?, Behavioral and Brain Sciences, 5, 01, (167), (1982).
  • , Does play matter? Functional and evolutionary aspects of animal and human play, Behavioral and Brain Sciences, 5, 01, (139), (1982).
  • , Selective Costs and Benefits in the Evolution of Learning, , 10.1016/S0065-3454(08)60046-7, (65-106), (1982).
  • , The flexibility and affective autonomy of play, Behavioral and Brain Sciences, 5, 01, (160), (1982).
  • , Explaining the evolutionary significance of intellectual play: Are we barking up the wrong tree?, Behavioral and Brain Sciences, 5, 01, (166), (1982).
  • , Play stimulated by environmental complexity alters the brain and improves learning abilities in rodents, primates, and possibly humans, Behavioral and Brain Sciences, 5, 01, (164), (1982).
  • , Why does play matter?, Behavioral and Brain Sciences, 5, 01, (169), (1982).
  • , Comparison matters: Curiosity, bears, surplus energy, and why reptiles do not play, Behavioral and Brain Sciences, 5, 01, (159), (1982).
  • , The facts about fantasy, Behavioral and Brain Sciences, 5, 01, (172), (1982).
  • , Skill and intelligence: The functions of play, Behavioral and Brain Sciences, 5, 01, (163), (1982).
  • , Functions of play: First steps toward evolutionary explanation, Behavioral and Brain Sciences, 5, 01, (157), (1982).
  • , Hypotheses about play, Behavioral and Brain Sciences, 5, 01, (158), (1982).
  • , The current state of play, Behavioral and Brain Sciences, 5, 01, (172), (1982).
  • , Toward an individualistic ontology for cultural evolution, Behavioral and Brain Sciences, 4, 02, (242), (1981).
  • , A fourth approach to the study of learning: Are “processes” really necessary?, Behavioral and Brain Sciences, 4, 01, (151), (1981).
  • , Is hippocampal theta an artifact?, Behavioral and Brain Sciences, 4, 03, (480), (1981).
  • , Needed: More data on the reticular information, Behavioral and Brain Sciences, 4, 03, (485), (1981).
  • , The logical relation between cultural and biological evolution: On to the next question, Behavioral and Brain Sciences, 4, 02, (235), (1981).
  • , Discussing learning: The quandary of substance, Behavioral and Brain Sciences, 4, 01, (146), (1981).
  • , Multiple-level evolution: A disagreement to disagree, Behavioral and Brain Sciences, 4, 02, (253), (1981).
  • , Can the decomposition of attention clarify some clinical issues?, Behavioral and Brain Sciences, 4, 03, (477), (1981).
  • , On constraints and adaptation, Behavioral and Brain Sciences, 4, 02, (244), (1981).
  • , A functional view of learning, Behavioral and Brain Sciences, 4, 01, (153), (1981).
  • , An atropine-sensitive and a less atropine-sensitive system, Behavioral and Brain Sciences, 4, 03, (493), (1981).
  • , Reticular formation, brain waves, and coma, Behavioral and Brain Sciences, 4, 03, (489), (1981).
  • , Behavioral problems related to the interpretation of brain rhythms, Behavioral and Brain Sciences, 4, 03, (477), (1981).
  • , Some problems with an “options” view of evolution, Behavioral and Brain Sciences, 4, 02, (241), (1981).
  • , The limits of natural selection, Behavioral and Brain Sciences, 4, 02, (244), (1981).
  • , Sociobiology and Levels of Explanation, American Anthropologist, 83, 1, (147-149), (2009).
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