Current and future plant invasions in protected areas: Does clonality matter?

Protected areas (PAs) play an important role in biodiversity conservation, but remain increasingly threatened by invasive alien plant species (IAPS) in conjunction with global climate change. The latter is modifying the distribution of the former, and the magnitude and direction of distributional changes are predicted to vary depending on species dispersal mode. Here, we address the question of whether clonality is expected to affect the future invasion pattern in PAs.

Because invasive alien species generally have broad physiological tolerances and/or specific traits that enhance their competitive performance or rapid adaptation to harsh environments, they may respond quickly to changing environmental conditions (Hoffmann & Sgro, 2011;Mathakutha et al., 2019;Warren et al., 2020;Whitney & Gabler, 2008). Therefore, it is a hot topic for ecologists and biological conservationists to explore the effects of climate change in biological invasions around the world.
Understanding how climate change affects the distribution of invasive alien species is of particular resonance in protected areas (PAs), which had been established to protect biodiversity, and threatened native species, habitats and ecosystems (Foxcroft et al., 2007(Foxcroft et al., , 2011(Foxcroft et al., , 2019. Climate change may further increase the capacity of alien species to invade PAs and subsequently damage the conservation efficiency of PAs (Foxcroft et al., 2007(Foxcroft et al., , 2011Gallardo et al., 2017;Padmanaba et al., 2017;Pěknicová & Berchová-Bímová, 2016). At the global scale, however, it is still unknown the mechanism on how climate change is expected to affect invasions in PAs.
The impact of climate change is recognized to vary according to life forms, generation times, reproduction modes and dispersal abilities in plants (Corlett & Westcott, 2013;Nicotra et al., 2010).
As a result, we can expect invasion risks in PAs to depend on whether invasive alien plant species (IAPS) are able to reproduce asexually or not (Gallardo et al., 2017;Gillson et al., 2013;Lamsal et al., 2018). Previous studies have shown that clonality can contribute greatly to plant invasions (e.g. Eckert et al., 2016;Fenollosa et al., 2016;Liu et al., 2006;Song et al., 2013). It has also been noted that many IAPS reproduce by clonal growth and that many of the most invasive plants in the world are clonal (Fenollosa et al., 2016;Liu et al., 2006;Yu et al., 2019). For instance, 2/3 of the most invasive plants in China and also about 2/3 of the world's worst invasive plants listed by the ISSG (Invasive Species Specialist Group) are clonal (Liu et al., 2006;Lowe et al., 2000). In addition to the ability to disperse by seeds, clonal plants can also spread their populations by clonal growth and may thus be less constrained by climate because they are not temperature-regulated regarding flowering and fruiting (Ye et al., 2014;Yu et al., 2019). Furthermore, clonal plants possess some distinguished characteristics that can assist them to quickly establish their populations in unexpectedly harsh environments (Negreiros et al., 2014). These differences may result in altered adaptability of clonal compared to non-clonal plants to environmental changes (Ye et al., 2014). Previous studies (e.g. Bellard et al., 2014;Burgess et al., 2017;Gillard et al., 2017;Osawa et al., 2019;Wan & Wang, 2018) used species distribution modellings (SDMs) to the distributions of these world's worst invasive plants. However, these studies only established correlative SDMs based on effects of environmental changes on IAPS distributions using presence and absence points. The early studies on SDMs do not allow to mechanistically model the direct effects of clonal versus non-clonal life strategies on IAPS distributions from local to global scales. Therefore, to develop adapted conservation strategies and reduce invasion risks, it is critical to know about the distributional responses of clonal versus non-clonal IAPS in PAs in the course of climate change.
The influence of climate change in the distribution of IAPS in PAs may also depend on biomes, that is the major vegetation complexes classified based on dominant vegetation types and associated climatic and other major environmental conditions (Bradley et al., 2010;Gallagher et al., 2010;Thuiller et al., 2005). Indeed, plant invasions differ greatly among different biomes because biotic and abiotic conditions vary considerably among them (Bradley et al., 2010;Gallagher et al., 2010). Furthermore, the abundance of clonal plants varies greatly among biomes (Kalusová et al., 2013;Rood et al., 2007). For instance, clonal plants are dominant species in grasslands, wetlands and tundra, but occur less frequently in conifer forests (Klimešová et al., 2017). Therefore, the influence of climate change in the prevalence of clonal and non-clonal IAPS in PAs can also differ among biomes. So far, however, no study has tested whether the susceptibility of PAs to clonal and non-clonal IAPS differs among biomes in the course of climate change.
We modelled the current and future distribution of 36 plant species found in the list of "100 of the world's worst invasive alien species" established by the Invasive Species Specialist Group (Lowe et al., 2000). We split this set of species into clonal and non-clonal categories and assessed their current probability to invade global PAs distributed in 16 biomes and seven realms as well as their future in- (c) Will this change be the same among clonal and non-clonal plants?

| Species data
The Invasive Species Specialist Group (ISSG) of the International Union for Conservation of Nature (IUCN) has compiled a list of "100 of world's worst invasive alien species" (Lowe et al., 2000; http:// www.issg.org/datab ase/speci es/search.asp?st=100ss). We used the 36 IAPS from this list (Table S1) as the most geographically and taxonomically representative set of the most noxious IAPS around the world, causing significant impacts on biodiversity and/or human activity. Clonal plants are those that reproduce asexually by means of vegetative offspring that remain attached to the parent, at least until they establish . We identified clonal IAPS based on whether the species has potential clonality in life-history strategies from the perspectives on the clonal and bud bank traits (Klimešová et al., 2017). First, we checked whether 36 species are clonal from the list of CLO-PLA3 database (www.clopla.butbn.cas.cz/). Then, we determined the clonal and bud bank traits for each species based on the TRY database (www.try-db.org/TryWe b/Home.php; Kattge et al., 2020) and the Botanical Information and Ecology Network (BIEN) database (Maitner et al., 2018). Finally, clonal plant species could be identified if the species was listed in CLO-PLA3 database and had the clonal and bud bank traits in life-history strategies.
Among the 36 IAPS, 13 were identified as non-clonal and 23 as clonal according to ISSG and other references (Liu et al., 2006; Table 1 and   Table S1). Contemporaneous occurrence data with geographic coordinates were obtained for each IAPS from several online databases including: (a) the Global Biodiversity Information Facility (GBIF; www. gbif.org), (b) LIFEMAPPER (www.lifem apper.com), (c) SPECIESLINK (www.splink.cria.org.br), (d) the Chinese Virtual Herbarium (CVH; www.cvh.org.cn), (e) the IUCN/SSC ISSG (Lowe et al., 2000) and (f) published literatures. All extracted occurrences were resampled at 2.5-arc-minute resolution (ca. 5 km at the equator), and duplicated records were removed to reduce the effect of sampling bias. Overall, we obtained 70,020 unique records, that is 1,945 records for each IAPS on average (ranging from 52 for the coralberry Ardisia elliptica to 26,506 for the purple loosestrife Lythrum salicaria) across the world, with the exception of the Sahara region, most regions of Russia, northern Canada and Greenland (Table S1 and Figure S1).

| Climate data
Nineteen climatic variables derived from the WorldClim database (representing 1950-2000 averages; Table S2; Hijmans et al., 2005; www.world clim.org) were used for modelling purposes. We selected these variables at a 2.5-arc-minute resolution because a finer resolution would cast a false sense of precision despite potentially giving higher accuracy scores (Ramirez-Villegas & Jarvis, 2010). Among these variables, we removed those with Pearson's correlation coefficient |r| > 0.7 to avoid multi-collinearity effects in the parameter estimates of species distribution models (Elith et al., 2011). The four resulting variables were annual mean temperature, temperature seasonality, precipitation of the driest month and precipitation of the wettest quarter.

| Protected areas, biomes and realms
A global map of PAs was obtained from the World Database on Protected Areas (WDPA; http://www.wdpa.org/). We excluded protected seascape or PAs lacking information on area coverage.
We also excluded PAs too small to be represented in one grid cell (<2.5 × 2.5 arc minutes). Finally, we used more than 20,000 PAs whose size ranged from 1 to 194,166 cells.
The terrestrial area of the globe was further classified into 16 biomes, representing the major global plant communities determined by temperature and precipitation ( Figure S2; Olson et al., 2001). The map of these biomes was obtained from http://maps.tnc.org/gis_ data.html#ERA as described by the World Wildlife Fund (WWF) and The Nature Conservancy (TNC; Olson et al., 2001). Based on gridded maps of PAs and biomes, we assigned each PA to one of the 16 biome types using a majority function. This allowed us to analyse the effect of the biome type on the distribution of clonal and non-clonal IAPS in PAs under future climate change. We also assigned PAs to the types of realms based on a global ecoregion map from http:// maps.tnc.org/gis_data.html#ERA as described by the World Wildlife Fund (WWF) and The Nature Conservancy (TNC; Olson et al., 2001) through a majority function ( Figure S2).

| Modelling approach and evaluation
We projected the current and future global potential distributions of the 36 IAPS based on contemporary occurrence localities and current and future climatic data. We used three species distribution models, that is general linear models (GLM; McCullagh & Nelder, 1989), general additive models (GAM; Hastie & Tibshirani, 1986) and Maxent (Phillips et al., 2006). GLM is considered to result in simple, GAM in moderately complex and MaxEnt in highly complex response shapes (Mainali et al., 2015). We set the regularization multiplier (beta) to 1.5 to produce a smooth and general response shape that stands for a biologically realistic behaviour in Maxent. The maximum number of background points was set to 10,000, and we used a 10-fold cross-validation approach to remove bias with respect to recorded occurrence points.
We evaluated the predictive precision of the species distribution models using the area under the curve (AUC) of the receiver operation characteristic (ROC). The AUC values range from 0 (systematically wrong) to 1.0 (highest predictive ability), while a value of 0.5 indicates a random model fit. The three models built for each species with values above 0.7 were considered useful in our study.
We averaged the results of SDM across GLM, GAM and MaxEnt for each IAPS, and AUC values of SDMs were higher than 0.7. However, AUC was insufficient for assessing the performance of Maxent modelling. Therefore, we used a binomial test based on the omission rate to evaluate the performance of Maxent modelling for the 36 IPS (Anderson et al., 2002(Anderson et al., , 2003. The omission rates of training and test occurrence records were calculated as the proportion of the sample points within grid cells that were predicted to yield the absences of the species for the occurrence localities of test data (Anderson et al., 2002(Anderson et al., , 2003. Then, one-sided p-values were used to test the null hypothesis, and the test points are predicted no better than those by a random prediction with the same fractional predicted area (Anderson et al., 2002). The binomial probabilities were based on 11 common threshold defaults by Maxent modelling (detailed information in Phillips et al., 2006). Although the training and test omission rates may not be sufficient, a low omission rate (i.e. 15%) is a necessary condition for a good model (Anderson et al., 2002, TA B L E 1 Change in the probability of invasive alien plant species to invade protected areas between the current situation and the high concentration scenario (RCP 8.5) according to the realm distribution of protected areas

| Potential of IAPS to invade PAs
We analysed the probability of clonal IAPS, non-clonal IAPS and all IAPS (clonal plus non-clonal) to invade PAs at three geographic levels (globe, biome and PA). To do so, we calculated the current and future potential distribution for each species, climate model and climate scenario.
To estimate the future distribution of single IAPS under the three concentration scenarios, we superimposed the potential future distribution maps of single IAPS for each of the 4 GCMs ×3 RCPs with identical weight. We then averaged the potential distribution of cooccurring IAPS in the low, medium and high greenhouse gas concentration scenarios and analysed the potential of co-occurring IAPS to colonize PAs using the present distributions as a basis for comparison. Many previous studies have set a presence/absence threshold for each individual species to estimate species richness through ensemble modelling. However, these thresholds are problematic and can produce bias in predictions (Calabrese et al., 2014). Here, we used the modified method of Calabrese et al. (2014) to compute the invasion extent of co-occurring IAPS in each pixel: where E j represents the current or future invasion extent of cooccurring IAPS in pixel j, k is the number of species in pixel j, and P j,k is the probability of potential distribution of species i in pixel j.
We calculated the probability of multiple IAPS to invade the PA as follows: where S t is the current or future probability of co-occurring IAPS to invade PA t, X j an indicator of the distribution possibility of co-occurring We calculated the change in the probability of multiple IAPS for each PA between the current scenario and the 2080s (in the low, medium and high concentration scenarios): where A i is the change in the probability of multiple IAPS to invade PAs and S Future and S Current are the future and current probabilities of multiple IAPS to invade PAs. We calculated the probability change for clonal IAPS, non-clonal IAPS and all IAPS.

| Risk hot spots of IAPS invasions in PAs
We used the Optimizing Hot Spot Analysis (ESRI, 2014)

| Influences of climate change and clonality on plant invasions in PAs
The future probability of all IAPS (i.e. clonal plus non-clonal IAPS) to invade PAs changed very little from the current situation to the low (−1.94%), medium (−1.40%) and the high greenhouse gas concentration scenarios (0.05%) (Figure 1). The consequence of climate change in the probability of clonal IAPS alone and non-clonal IAPS alone to invade PAs was also small (probability change between present and future was less than ±5%; Figure 1 Figure 2). However, this probability change was much smaller for clonal IAPS than for nonclonal IAPS in Tundra (21.14 vs. 68.42%; Figure 2). Over Rock and Ice, the change was slightly positive for clonal IAPS (11.56%), but negative for non-clonal IAPS (−39.81%; Figure 2). Clonality had little impact on the probability change in the other biomes ( Figure 2).
The largest impacts of climate change in the invasion probability of both clonal and non-clonal IAPS occur in PAs of Nearctic and Palearctic (

| Hot spots of plant invasions
Based on the distribution of all IAPS, invasion hot spots were similar under the current and future climate scenarios ( Figure S4). They

| Effects of climate change in plant invasions in PAs
At the global scale, climate change had little impact on the probability of IAPS to invade PAs, suggesting that global climate change will unlikely promote the invasions of our set of IAPS into PAs across the globe. This finding is consistent with that of a recent study showing that the potential distributions of species, including plants, animals and microbes, were not significantly related to global climate change (e.g. Bellard et al., 2013Bellard et al., , 2014. However, when we analysed F I G U R E 1 Change in the probability of clonal, non-clonal and all (clonal plus non-clonal) invasive alien plant species to invade global protected areas between the current situation and the three future concentration scenarios (low, medium and high). Dash lines represent values of +5% and −5%. RCPs 2.6, 4.5 and 8.5 were used for the low, medium and high climate scenarios. The probabilities were derived from assemble species distribution modelling for invasive alien plant species On the other hand, climate change was predicted to decrease the range of IAPS in PAs in five biomes mainly located in tropical F I G U R E 3 Hot spots of probability change of all (clonal plus non-clonal), clonal and non-clonal invasive alien plant species to invade protected areas in the current situation and the high gas concentration scenario (RCP 8.5). The probabilities and hot spots were derived from assemble species distribution modelling. The colour from blue to red represented the increasing probability of invasive alien plant species to invade protected areas and subtropical climates. Global warming is expected to reduce plant diversity in tropical areas, and IAPS would be no exception (Bellard et al., 2014;Brodie et al., 2012). Hence, regarding the limited financial resources available for coordinated regional conservation actions, we believe fewer efforts can be spent in PAs located in these biomes.

| Impacts of clonality on plant invasions in PAs
While clonality had little impact on the invasion risk in PAs medi- Aquatic ecosystems are prone to biological invasions, and many inland aquatic ecosystems in the world are heavily invaded by aquatic clonal plants (Eckert et al., 2016;Hussner et al., 2017;Santamaría, 2002;Teixeira et al., 2017). For some aquatics such as the common water hyacinth Eichhornia crassipes, the main way to spread and invade is by clonal growth, and the spread of clonal propagules is also much easier in such ecosystems (Herben & Klimešová, 2020;Yu et al., 2019). Hence, in the future, we need to pay much attention to clonal IAPS in PAs which function to conserve Inland Water.
Clonal plant species play an important role in Temperate grasslands, Savannas and Shrublands, and PAs found in this biome usually harbour a rich biodiversity (Olson et al., 2001). Clonal plants are able to successfully invade new habitats because they do not necessarily need to establish a population by producing seeds and they just need a single individual to do well enough to produce ramets (Bittebiere et al., 2020;Byun et al., 2015). Furthermore, clonal plants are widely distributed in Temperate Grasslands, Savannas and Shrublands and are sensitive to increasing nitrogen deposition (Negreiros et al., 2014;Osborne et al., 2018). Enhanced nitrogen deposition may increase the number of ramets of IAPS, which could lead to dynamic changes in plant communities in PAs of the aforementioned biomes (Negreiros et al., 2014;Osborne et al., 2018).  (Negreiros et al., 2014). Efforts should then be allocated mostly to clonal plants and not to non-clonal plants (Goldberg et al., 2020;van Kleunen et al., 2001;Kleyer & Minden, 2015). In Tundra, future climate change will probably increase the invasions of non-clonal IAPS in PAs much more than that of clonal IAPS. Hence, we need to pay attention to the invasion of non-clonal IAPS in PAs of this biome.

| Current and future hot spots of plant invasions
We found that hot spots of the 36 worst IAPS under all three future climate scenarios matched with current hot spots. We stressed the importance of monitoring PAs in regions such as southwestern and southeastern Australia, New Zealand, Mexico, southeastern Asia and southern China, which are also known to be biodiversity hot spots of conservation priorities (www.conse rvati on.org/how/ pages/ hotsp ots.aspx; Myers et al., 2000). The overlap between invasion hot spots and biodiversity hot spots stands for a serious problem as the expansion of IAPS, facilitated or unfacilitated by climate change, will decrease the space available for native species, which is likely to lead to ecosystem disorders and, ultimately, to species extinctions (Bellard et al., 2013(Bellard et al., , 2014. In some regions, IAPS are projected to spread from one into other PAs (Foxcroft et al., 2011(Foxcroft et al., , 2017(Foxcroft et al., , 2019. For example, the invasion hot spots showed a tendency of moving northward in Europe, and the density of invasion hot spots in northern Latin America is higher in the future than today based on our results. Rapid globalization associated with high human mobility promotes the establishment of populations of IAPS in new habitats (Chapman et al., 2017;van Kleunen et al., 2020). For example, international trade is a critical force for the spread of IAPS due to frequent escapes and releases of introduced species into the wild (Chapman et al., 2017;Seebens et al., 2015). Furthermore, the economic use of IAPS plays a significant role in their naturalization success (van Kleunen et al., 2020). Perhaps the highest naturalization success for IAPS is its use as animal food or its use in horticulture or as ornamentals (van Kleunen et al., 2020). Invasion patterns are governed to a large extent by the global trade networks connecting source areas of IAPS and their dispersal through multiple networks (e.g. trade and transport; Chapman et al., 2017;Seebens et al., 2015). Our results do not explicitly address these invasion pathways, but they provide spatially explicit information about invasion hot spots around the world.
Therefore, rapid globalization and high human mobility, coupled with distributional changes, could promote plant invasions in global PAs under climate change (Foxcroft et al., 2017;Seebens et al., 2015).
When observing invasions of clonal IAPS in a PA, we need to take immediate measures to prevent the spread of clonal IAPS, thus avoiding to "infect" other PAs around the invaded region. These measures include developing global indicators of biological invasions and designing long-term management plans at different geographical scales (Foxcroft et al., 2017). These measures should not be taken in a hurry, and it is important to commit to scientific assessments such as the species distribution and life history of clonal species (Herben et al., 2014;Thuiller et al., 2012). Resource utilization strategies of IAPS could promote their invasions (Funk & Vitousek, 2007;Parepa et al., 2013). IAPS must have access to available resources (e.g. nutrients, light, and water) to successfully invade a community and will have a high chance of invasion success if they do not encounter intense competition for these resources from resident species (Davis et al., 2000;Parepa et al., 2013). High growth rate and the ability to rapidly exploit available resources (e.g. nitrogen nutrients) are widely recognized as fundamental plant strategies and are a potential determinant of invasion success (Davis et al., 2000;Funk & Vitousek, 2007;Parepa et al., 2013). Therefore, nitrogen deposition can promote growth and provide eco-physiological advantages for IAPS (Bradley et al., 2010;Funk & Vitousek, 2007;Perry et al., 2010). Resources such as nutrients, light and water taken up by plants can be easily released into soils through hydraulic redistribution and can also be translocated by clonal integration within a plant clonal network Ye et al., 2016). Clonal IAPS can benefit from high resource availability through clonal integration (Song et al., 2013;Wang et al., 2017;Yu et al., 2019). When detecting the distribution of IAPS, especially in invasion hot spots, areas of high resource availability (e.g. those with high nitrogen depositions) should receive special attention in strategies to prevent and control invasions of IAPS under climate change (Gough et al., 2012). However, early remediation actions have shown to be more effective and less costly than measures that are taken only after massive invasion success in North America, New Zealand and Europe, although details about the exact strategy related to the timing, frequency and intensity of actions tend to be species-specific (Foxcroft et al., 2011;Meier et al., 2014).
IAPS can invade PAs, benefiting from clonal reproduction and plasticity (Fenollosa et al., 2016). Many plants have the capacity for facultative clonal growth Liu et al., 2006;Song et al., 2013), and clonal plants in general have the capacity for facultative sexual reproduction Klimešová et al., 2017).
Usually, PAs have rich species diversity, which may resist to plant invasion (Crutsinger et al., 2008;Dalrymple et al., 2015;Maron & Marler, 2007). Clonal IAPS can switch strategies between sexual and non-sexual reproductions for shaping species coexistence so that they can adapt to different levels of species diversity and climate change (Yamamichi et al., 2020;Zobel, 2008). Asexual plants change just as often and just as fast as do sexual plants when introduced to a new range (Dalrymple et al., 2015). Furthermore, clonal plasticity facilitates the adaptation of IAPS to rapid changing environments (Nicotra et al., 2010;Wang et al., 2018). Clonal plasticity could enhance the exploitation of resource heterogeneity by clonal IAPS, which have a significant contribution to maintenance or improvement of fitness under climate change (Nicotra et al., 2010;Santamaría, 2002;Wang et al., 2021). Clonal reproduction and plasticity may make the difference in invasion ability to PAs between clonal and non-clonal IAPS under climate change. Thus, clonality may be a key indicator of IAPS to invade PAs under climate change around the world.
For targeted observations of clonal IAPS, we suggest using Figure 3 to facilitate negotiations with stakeholders and decisionmakers. In Figure 3, (Drenovsky et al., 2012). Variation of these traits may affect the distribution pattern and species interactions of clonal and non-clonal plants under environmental changes (Bittebiere et al., 2019;Herben & Klimešová, 2020) and thus the invasion success of clonal versus non-clonal IAPS (Wang et al., 2017).
Hence, other functional traits of IAPS than clonality could be integrated into species distribution models to improve their performance at the global scale (Benito Garzón et al., 2019). In addition, dynamic hybrid models combined with species distribution models may facilitate the development of optimization strategies (Buchadas et al., 2017).
These tools are essential for designing long-term management plans at the national to regional scales in order to create a concerted mitigation strategy for IAPS invasions into PAs under climate change.

| Limitations
Although our study provided the global maps of current and future plant invasions in protected areas for clonal and non-clonal plants,  (Gough et al., 2012;Liu et al., 2006;Wang et al., 2017). Third, we did not consider biome transition zones or in periods of biome transition.
In the transition biomes and realms, instability and heterogeneity can promote plant invasions. Finally, there are many uncertainties on SDM results (e.g. model transferability) for projecting distributions of IAPS across different spatial scales (Araújo et al., 2019;Buisson et al., 2010;Chen et al., 2019;Guo et al., 2015;Liu et al., 2020;Zurell et al., 2020). The reliability of transferring SDMs to new ranges and future climates has been widely debated (Liu et al., 2020). Model transferability is intrinsically determined by the significant relationships between environmental predictors and species distributions, and the number of occurrence records for modelling distributions of IAPS (Liu et al., 2020;Petitpierre et al., 2012). Our study only considered the relationships between climatic predictors and distributions of IAPS at the global scale. Future studies should take the relationships between other environmental predictors (e.g. land use and land cover, and soil factors) and distributions of IAPS into SDMs.
It is also important to collect a larger number of occurrence records as the input of SDMs for modelling distributions of IAPS (Araújo et al., 2019;Chen et al., 2019;Liu et al., 2020;Zurell et al., 2020).

| CON CLUS IONS
Global climate change may not promote the invasions of IAPS in PAs and plant clonality shows little impact at the global scale. However, climate change can markedly change plant invasion patterns in PAs at the scale of biomes and realms, and clonal and non-clonal plants also play contrasting roles in different biomes and realms. Therefore, to design effective strategies to prevent and control IAPS in PAs, biomes and plant reproductive traits should be carefully considered.

ACK N OWLED G EM ENTS
We thank Prof. Hai-Ning Qing for allowing us to use the CVH data.

CO N FLI C T O F I NTE R E S T
The authors have no interest or relationship, financial or otherwise that might be perceived as influencing the author's objectivity with this work and thus have no conflicts of interest to declare.

PE E R R E V I E W
The peer review history for this article is available at https://publo ns.com/publo n/10.1111/ddi.13425.

DATA AVA I L A B I L I T Y S TAT E M E N T
All of the data in this paper are downloaded from publicly accessi-